Kearn, G.C. (1976)
Observations on monogenean parasites from the nasal fossae of European rays: Empruthotrema raiae (MacCallum, 1916), Johnston and Tiegs, 1922 and E. torpedinis sp. nov. from Torpedo marmorata. Proceedings of the Institute of Biology and Pedology, Far–East Science Centre, Vladivostock, USSR. 34: 45–54.(In Russian, with English summary).
Chisholm, L.A. & Whittington, I.D. (1999)
A revision of the Merizocotylinae Johnston and Tiegs, 1922 (Monogenea: Monocotylidae) with descriptions of the new species of Empruthotrema Johnston and Tiegs, 1922 and Merizocotyle Cerfontaine, 1894. Journal of Natural History, 33(1), 1–28
DOI: 10.1080/002229399300452