NEWSLETTER 07/2012 28. July 2012



  • Corsica - Mediterranean Shark Research Group (Homepage)
  • Pr. Jean-Lou Justine, UMR 7138 Systematique, Adaptation, Evolution, Museum National d’Histoire Naturelle, Paris, France
  • David Jacoby, The Marine Biological Association of the UK, Plymouth, UK


Partner in Google-Maps






Many thanks to Dr. José De La Cruz - Agüero, Professor and Curator of CI, Centro Interdisciplinario de Ciencias Marinas (CICIMAR-IPN) for the permission to use images of:

Aetobatus narinari
Carcharhinus limbatus
Carcharhinus porosus
Cephaloscyllium ventriosum
Dasyatis brevis
Diplobatis ommata
Ginglymostoma cirratum
Gymnura crebripunctata
Gymnura marmorata
Heterodontus francisci
Heterodontus mexicanus
Hydrolagus colliei
Mustelus californicus
Mustelus henlei
Mustelus lunulatus
Myliobatis californica
Myliobatis longirostris
Narcine entemedor
Nasolamia velox
Parmaturus xaniurus
Platyrhinoidis triseriata
Raja equatorialis
Raja inornata
Rhinobatos glaucostigma
Rhinobatos leucorhynchus
Rhinobatos productus
Rhinoptera steindachneri
Rhizoprionodon longurio
Sphyrna lewini
Sphyrna zygaena
Squalus acanthias
Squatina californica
Triakis semifasciata
Urobatis concentricus
Urobatis halleri
Urobatis maculatus
Urotrygon aspidura
Urotrygon munda
Urotrygon chilensis
Urotrygon rogersi
Zapteryx exasperata







Thanks to 

Carlos Bustamente, Queensland, Australia for the images of Amblyraja frerichsiDipturus trachyderma and Zearaja chilensis






Chris Avila, Toronto for images of juvenile Atelomycterus baliensis


Simon Weigmann, Hamburg for images of Carcharhinus dussumieriChiloscyllium griseumChiloscyllium punctatumHemigaleus microstomaParagaleus randalli and Sphyrna lewini, e.g.:



Prof. Jean-Lou Justine, Paris for images of Hexanchus nakamuraiCarcharhinus brevipinna and Pastinachus sephen, e.g.:



Dr. Csilla Ari for images of Manta birostris, e.g.:



Please visit the species descriptions to see all new images!


Please send your images to shark-references!



New information at -  Host-Parasites-List (in preperation)



parasites list of Aetobatus narinari


Monogenea Cestoda
  • Acanthobothrium aetiobatis (SHIPLEY, 1900) [16175]
  • Acanthobothrium arlenae CAMPBELL & BEVERIDGE, 2002 [16128]
  • Acanthobothrium nicoyaense BROOKS & McCORQUODALE, 1995 [7562]
  • Acanthobothrium monski MARGUEE, BROOKS & BARRIGA, 1997 [7617]
  • Acanthobothrium tortum (LINTON, 1916) BAER & EUZET, 1962 (synonyms Onchobothrium tortum LINTON, 1916 [16172]) [16175] [16178]
  • Didymorhynchus southwelli, BEVERIDGE & CAMPBELL, 1988 [16112] [16255]
  • Dollfusiella aetobati (BEVERIDGE, 1990) [16112]
  • Eutetrarhynchus aetobati BEVERIDGE, 1990 [16252]
  • Hornellobothrium cobraformis SHIPLEY & HORNELL, 1906 [13130]
  • Kotorella pronosoma (STOSSICH, 1901) [16112]
  • Oncomegas australiensi TOTH, CAMPBELL & SCHMIDT, 1992 [16112] [15474] [16428]
  • Oncomegas sp. [15474]
  • Parachristianella baverstocki BEVERIDGE, 1990 [16112]
  • Proemotobothrium linstowi (SOUTHWELL, 1912) [16112]
  • Shirleyrhynchus aetobatidis (SHIPLEY & HORNELL, 1906) [16112] [16256]
  • Trimacracanthus aetobatidis (ROBINSON, 1959) [16254]
  • Trygonicola macroporus (SHIPLEY & HORNELL, 1906) [16112]
  • Tylocephalum aurangabadensis JADHAV & SHINDE, 1987 [16330]
  • Tylocephalum girindrai PRAMANIK & MANNA, 2007 [16377]
  • Echinocephalus sinensis KO, 1975 [7404]
  • Echinocephalus overstreeti DEARDORFF & KO, 1983 [7404]
  • Terranova scoliodontis (BAYLIS, 1931) [7404]
  • Eudactylina hornbosteli DEETS, 1994 [15305]



Please inform me about missing data!



Please support shark-references:

Shark-References would kindly like to ask you for your contribution to this project.

You have fulltext access to one of the following journals?


African Journal of Marine Science

African Natural History

South African Journal of Marine Science

South African Journal of Science

Please send me some of this missing papers (only for my use, not for download!)

Thank you very much for your support!


Many thanks for sending missing papers:

France Charles F. Cotton

Oliver Landemaine  







The KwaZulu-Natal Sharks Board is pleased to announce that it will be hosting the second SharksInternational Symposium in Durban, South Africa in early June 2014. 


Sharks International in Cairns, Australia in June 2010 was the first fully international elasmobranch conference in nearly 20 years.   It was highly successful, so much so that the overwhelming consensus among the delegates was that it should become a regular event. 


We believe that a four-year interval between major conferences allows researchers sufficient time to assemble new findings to share with their peers and to budget for what promises to be another memorable meeting.


The structure of the 2014 conference will be identical to the Cairns meeting.  There will four full days of presentations, commencing Monday 2 June and ending on Friday 6 June, with a rest day on Wednesday to allow delegates to relax or to sample the many tourist delights of Durban and surrounds.   Posters will be welcome and parallel sessions will be avoided if possible.   The meeting will end with a gala dinner on Friday.


We have been promised a generous grant from the Department of Economic Development and Tourism of the KwaZulu-Natal provincial government.  This will allow us to keep registrations fees low and to offer travel grants to worthy students and young researchers.  We are also hoping that it will enable us to sponsor some keynote speakers.   Durban and nearby Umhlanga has world class hotel-based conference venues and we are having a tricky task choosing the most suitable one.  


Durban is a vibrant city on the east coast of South Africa, with a blend of Zulu, Western and Indian cultures.  The climate is mild; the average June temperature range is 13-23°C.  It has world class beaches and is within easy reach of the Drakensberg mountains and some of the finest game parks in Africa.  We hope that elasmobranch researchers will not only take this opportunity to interact with colleagues at Sharks International but will enjoy the many tourist attractions of the region.


For the marathon runners, I would urge you to enter the Comrades Marathon, one of the world’s top ultra-marathons, run annually between Pietermaritzburg and Durban.  It is a mere 89 km and is scheduled for Sunday 1 June 2014.


We are in the process of developing a conference website which will include details of some tour options for the rest day and more extended tours after the conference.  We will let you know when it is up and running.  In the meantime you can contact us on       


Please make Sharks International, Durban a priority in June 2014.




Geremy Cliff

Head - Research
KwaZulu-Natal Sharks Board
Private Bag 2

Umhlanga Rocks,4320

South Africa


sharks board


More Meetings see:


Now freely downloadable online!

Selachians from the Upper Cretaceous (Santonian) Hosta Tongue of the Point Lookout Sandstone, central New Mexico

by Jim Bourdon, Keith Wright, Spencer G. Lucas, Justin A. Spielmann and Randy Pence

Thanks to Spencer G. Lucas, New Mexico Museum of Natural History and Science for the link!


New Paper




AJEMIAN, M.J. & POWERS, S.P. & MURDOCH, T.J.T. 2012 Estimating the Potential Impacts of Large Mesopredators on Benthic Resources: Integrative Assessment of Spotted Eagle Ray Foraging Ecology in Bermuda. PLoS ONE, 7 (7): e40227

ANDERSON, E.T. & FRASCA, S. & ASAKAWA, M.G. & FATZINGER, M.H. & JOHNSON, J. & MARCHETERE, K. & GOODALE, L. & RISATTI, G.R. & HARMS, C.A. 2012 Splenic mycobacteriosis in an Atlantic guitarfish, Rhinobatos lentiginosus Garman. Journal of Fish Diseases, 35 (7):  541-544

ARI, C. 2012 Visual abilities and social interaction of manta rays with the largest brain of fishes and the possible underlying neurological structures. Abstract. In: Abstracts of the International Behavioral Neuroscience Society, Kailua-Kona, Hawaii, USA, June 5-10, 2012, 21: 47

BARNETT, A. & ABRANTES, K.G. & SEYMOUR, J. & FITZPATRICK, R. 2012 Residency and Spatial Use by Reef Sharks of an Isolated Seamount and Its Implications for Conservation. PLoS ONE, 7 (5): e36574

BARNETT, A. & SEMMENS, J.M. 2012 Sequential movement into coastal habitats and high spatial overlap of predator and prey suggest high predation pressure in protected areas. Oikos, 121 (6): 882-890

BENDIAB, A.A.T. & MOUFFOK, S. & BOUTIBA, Z. 2012 Reproductive biology and growth of Lesser Spotted Dogfish Scyliorhinus canicula (Linnaeus, 1758) in Western Algerian coasts (Chondrichthyes, Scyliorhinidae). Biodiversity Journal, 3 (1): 41-48

BEVERIDGE, I. & CAMPBELL, R.A. 2012 Bathygrillotia n. g. (Cestoda: Trypanorhyncha), with redescriptions of B. rowei (Campbell, 1977) n. comb. and B. kovalevae (Palm, 1995) n. comb. Systematic Parasitology, 82 (3): 249-259

BOUGATEF, A. & BALTI, R. & SILA, A. & NASRI, R. & GRAIAA, G. & NASRI, M. 2012 Recovery and physicochemical properties of smooth hound (Mustelus mustelus) skin gelatin. Food Science and Technology, 48 (2): 248-254

BUSTAMANTE, C. & LAMILLA, J. & CONCHA, F. & EBERT, D.A. & BENNETT, M.B. 2012 Morphological Characters of the Thickbody Skate Amblyraja frerichsi (Krefft 1968) (Rajiformes: Rajidae), with Notes on Its Biology. PLoS ONE 7(6): e39963

CHIN, A. & TOBIN, A. & SIMPFENDORFER, C. & HEUPEL, M.  2012 Reef sharks and inshore habitats: patterns of occurrence and implications for vulnerability. Marine Ecology Progress Series, 460: 115-125

CLARKE, C. & LEA, J. & ORMOND, R. 2012 Comparative abundance of reef sharks in the Western Indian Ocean. Proceedings of the 12th International Coral Reef Symposium, Cairns, Australia, 9-13 July 2012: 5 pp

CONCHA, F. & ODDONE, M.C. & BUSTAMANTE, C. & MORALES, N. 2012 Egg capsules of the yellownose skate Zearaja chilensis (Guichenot 1848) and the roughskin skate Dipturus trachyderma (Krefft and Stehmann 1974) (Rajiformes: Rajidae) from the south-eastern Pacific Ocean. Ichthyological Research, in press

CONDON, N.E. & FRIEDMAN, J.R. & DRAZEN, J.C. 2012 Metabolic enzyme activities in shallow- and deep-water chondrichthyans: implications for metabolic and locomotor capacity. Marine Biology, 159 (8):

CRESSEY, D. 2012 Shark species more diverse than thought. Nature News, online

DE LA CRUZ-AGÜERO, J. & GARCÍA-RODRÍGUEZ, F.J. & COTA-GÓMEZ, V.M. & MELO-BARRERA, F.N. & GONZÁLEZ-ARMAS, R. 2012 Morphometric and molecular data on two mitochondrial genes of a newly discovered chimaeran fish (Hydrolagus melanophasma, Chondrichthyes). Ocean Science Journal, 47 (2): 147-153

DELANEY, D.G. & JOHNSON, R. & BESTER, M.N. & GENNARI, E. 2012 Accuracy of Using Visual Identification of White Sharks to Estimate Residency Patterns. PLoS ONE, 7 (4): e34753

DELI ANTONI, M.Y. & DELPIANI, G.E. & DELPIANI, S.M. & MABRAGAÑA, E. & DÍAZ DE ASTARLOA, J.M. 2012 An aberrant extra fin in Zearaja chilensis (Chondrichthyes: Rajidae). Cybium,36 (2): 403-405

DUDGEON, C.L. & BLOWER, D.C. & BRODERICK, D. & GILES, J.L. & HOLMES, B.J. & KASHIWAGI, T. & KRÜCK, N.C. & MORGAN, J.A.T. & TILLETT, B.J. & OVENDEN, J.R. 2012 Corrigendum to "A review of the application of molecular genetics for fisheries management and conservation of sharks and rays. (vol 80, pg 1789, 2012)" Journal of Fish Biology, 80 (7): 2649

FERREIRA, L.C. & VOOREN, C.M. 2012 Diet of the lesser electric ray Narcine brasiliensis (Olfers, 1831) (Elasmobranchii, Narcinidae) in southern Brazil. Pan-American Journal of Aquatic Sciences, 7 (1): 37-44

FISHERIES AND OCEANS CANADA 2012 Canada’s Progress Report on the Implementation of Key Actions Taken Pursuant to the National Plan of Action on the Conservation and Management of Sharks (March 2007).

GARIBALDI, F. & ORSI RELINI, L. 2012 Record of Carcharhinus falciformis (Bibron in Müller & Henle, 1839), in Italian waters (Ligurian Sea, Northwestern Mediterranean). Cybium,36 (2): 399-400

GRIJALBA-BENDECK, M. & POLO-SILVA, C. & ACEVEDO, K. & MORENO, F. & MOJICA, D. 2012 Aspectos tróficos y reproductivos de algunos batoideos capturados en Santa Marta, Mar Caribe de Colombia. Latin American Journal of Aquatic Research, 40 (2): 300 -315

GUSTAFSSON, O.S.E. & EKSTRÖM, P. & KRÖGER, R.H.H. 2012 Sturgeons, sharks, and rays have multifocal crystalline lenses and similar lens suspension apparatuses. Journal of Morphology, 273 (7): 746-753

HASELI, M. & MALEK, M. & PALM, H.W. & IVANOV, V.A. 2012 Two new species of Echinobothrium van Beneden, 1849 (Cestoda: Diphyllidea) from the Persian Gulf. Systematic Parasitology, 82 (3): 201-209

JACOBY, D.M.P. & BROOKS, E.J. & CROFT, D.P. & SIMS, D.W. 2012 Developing a deeper understanding of animal movements and spatial dynamics through novel application of network analyses. Methods in Ecology and Evolution, 3 (3): 574-583

KASHIWAGI, T. & BRODERICK, D. & LANCE, S.L. & BENNETT, M.B. & OVENDEN, J.R. 2012 Development and characterization of ten microsatellite loci for the reef manta ray Manta alfredi. Conservation Genetics Resources, in press

KING, J.R. & MCPHIE, R.P. & MORRISON, P.R.  2012 Biological results of the Strait of Georgia spiny dogfish (Squalus suckleyi) longline survey October 7-15, 2011.  Canadian Technical Report of Fisheries and Aquatic Sciences, 2975: 24 pp

LE PORT, A. & LAVERY, S. & MONTGOMERY, J.C. 2012 Conservation of coastal stingrays: seasonal abundance and population structure of the short-tailed stingray Dasyatis brevicaudata at a Marine Protected Area. ICES Journal of Marine Science, in press

LECLERC, L.-M.E. & LYDERSEN, C. & HAUG, T. & BACHMANN, L. & FISK, A.T. & KOVACS, K.M. 2012 A missing piece in the Arctic food web puzzle? Stomach contents of Greenland sharks sampled in Svalbard, Norway. Polar Biology, 35 (8): 1197-1208

LOPEZ, K.S. & BARRÍA, M.P. & MELÉNDEZ, C.R. 2012 Feeding and trophic relationships of two highly migratory sharks in the eastern south Pacific Ocean. Pan-American Journal of Aquatic Sciences, 7 (1): 50-56

MARQUES, F.P.L. & JENSEN, K. & CAIRA, J.N. 2012 Ahamulina n. gen. (Cestoda: Diphyllidea) from the polkadot catshark, Scyliorhinus besnardi (Carcharhiniformes: Scyliorhinidae), off Brazil. Zootaxa, 3352: 51-59

MCKINNEY, J.A. & HOFFMAYER, E.R. & WU, W. & FULFORD, R. & HENDON, J. 2012 Feeding habitat of the whale shark Rhincodon typus in the northern Gulf of Mexico determined using species distribution modelling. Marine Ecology Progress Series, 458: 199-211

MENDONÇA, S.A. & MACENA, B.C.L. & CREIO, E. & VIANA, D.L. & VIANA, D.F. & HAZIN, F.H.V. 2012 Record of a pregnant Mobula thurstoni and occurrence of Manta birostris Myliobatiformes: Mobulidae) in the vicinity of Saint Peter and Saint Paul Archipelago (Equatorial Atlantic). Pan-American Journal of Aquatic Sciences, 7 (1): 21-26

MEREDITH, T.L. & CAPRIO, J. & KAJIURA, S.M. 2012 Sensitivity and specificity of the olfactory epithelia of two elasmobranch species to bile salts. Journal of Experimental Biology, 215: 2660-2667

MONDO, K. & HAMMERSCHLAG, N. & BASILE, M. & PABLO, J. & BANACK, S.A. & MASH, D.C. 2012 Cyanobacterial neurotoxin beta-N-methylamino-L-alanine (BMAA) in shark fins. Marine Drugs, 10 (2): 509-520

NEFF, C.L. & YANG, J.Y.H. 2012 Shark bites and public attitudes: Policy implications from the first before and after shark bite survey. Marine Policy, in press

PARAMO, J. & WOLFF, M. & SAINT-PAUL, U. 2012 Deep-sea fish assemblages in the Colombian Caribbean Sea. Fisheries Research, 125-126: 87-98

PINHAL, D. & SHIVJI, M.S. & NACHTIGALL, P.G. & CHAPMAN, D.D. & MARTINS, C. 2012 A Streamlined DNA Tool for Global Identification of Heavily Exploited Coastal Shark Species (Genus Rhizoprionodon). PLoS ONE, 7 (4): e34797

QUEIROLO, D. & GAETE, E. & MONTENEGRO, I. & SORIGUER, M.C. & ERZINI, K.  2012 Behaviour of fish by-catch in the mouth of a crustacean trawl. Journal of Fish Biology, 80 (7): 2517-2527

RABEHAGASOA, N. & LORRAIN, A. & BACH, P. & POTIER, M. & JAQUEMET, S. & RICHARD, P. & MÉNARD, F. 2012 Isotopic niches of the blue shark Prionace glauca and the silky shark Carcharhinus falciformis in the southwestern Indian Ocean. Endangered Species Research, 17 (1): 83-92

RAMÍREZ-MOSQUEDA, E. & PÉREZ-JIMÉNEZ, J.C. & MENDOZA-CARRANZA, M. 2012 Reproductive parameters of the southern stingray Dasyatis americana in southern gulf of Mexico. Latin American Journal of Aquatic Research, 40 (2): 335-344

RENWART, M. & MALLEFET, J. 2012 Light emitting system in a deep sea shark: Etmopterus spinax (Squaloidea: Etmopteridae). Abstract. Luminescence, 27 (2): 154

ROGERS, P.J. & HUVENEERS, C. & PAGE, B. & HAMER, D.J. & GOLDSWORTHY, S.D. & MITCHELL, J.G. & SEURONT, L.A. 2012 A quantitative comparison of the diets of sympatric pelagic sharks in gulf and shelf ecosystems off southern Australia. ICES Journal of Marine Science, in press

SÁEZ, S. & LAMILLA, J. 2012 Claves taxonómicas para el reconocimiento dentario en taxa del superorden Rajomorphii de Chile (Chondrichthyes, Batoidea) (Taxonomic dental keys for the Chilean taxa of the superorder
Rajomorphii (Chondrichthyes, Batoidea)). Latin American Journal of Aquatic Research, 40 (2): 282-291

SÉRET, B. & LAST, P.R.  2012 New deep water skates of the genus Notoraja Ishiyama, 1958 (Rajoidei, Arhynchobatidae) from the southwest Pacific. Zoosystema, 34 (2): 319-341

SOLIMAN, V.S. & BURCE, R.B. 2012 Marine park with whale shark and adjacent to sardine fishery. Proceedings of the 12th International Coral Reef Symposium, Cairns, Australia, 9-13 July 2012: 5 pp

SPERONE, E. & PARISE, G. & LEONE, A. & MILAZZO, C. & CIRCOSTA, V. & SANTORO, G. & PAOLILLO, G. & MICARELLI, P. & TRIPEPI, S. 2012 Spatiotemporal patterns of distribution of large predatory sharks in Calabria (central Mediterranean, southern Italy). ACTA ADRIATICA, 53 (1): 13-24

TAGLIAFICO, A. & RAGO, N. & RANGEL, S. & MENDOZA, J. 2012 Exploitation and reproduction of the spotted eagle ray (Aetobatus narinari) in the Los Frailes Archipelago, Venezuela. Fishery Bulletin, 110 (3): 307- 316

THEODOSIOU, N.A. & SIMEONE, A. 2012 Evidence of a rudimentary colon in the elasmobranch, Leucoraja erinacea. Development Genes and Evolution, 222 (4): 237-243

TOMITA, T. & TODA, M. & UEDA, K. & UCHIDA, S. & NAKAYA, K. 2012 Live-bearing manta ray: how the embryo acquires oxygen without placenta and umbilical cord. Biology Letters, in press

USHAN, M. & WOOD, E. & SALEEM, M. & SATTAR, S.A. 2012 Maldives Sharkwatch Report for 2009 – 2010. Proceedings of the 12th International Coral Reef Symposium, Cairns, Australia, 9-13 July 2012: 4 pp

WATANABE, Y.Y. & LYDERSEN, C. & FISK, A.T. & KOVACS, K.M. 2012 The slowest fish: Swim speed and tail-beat frequency of Greenland sharks. Journal of Experimental Marine Biology and Ecology, 426-427: 5-11

WITT, M.J. & HARDY, T. & JOHNSON, L. & MCCLELLAN, C.M. & PIKESLEY, S.K. & RANGER, S. & RICHARDSON, P.B. & SOLANDT, J.-L. & SPEEDIE, C. & WILLIAMS, R. & GODLEY, B.J. 2012 Basking sharks in the northeast Atlantic: spatio-temporal trends from sightings in UK waters. Marine Ecology Progress Series, 459: 121-134

WUERINGER, B.E. & JNR, L.S. & KAJIURA, S.M. & TIBBETTS, I.R. & HART, N.S. & COLLIN, S.P. 2012 Electric Field Detection in Sawfish and Shovelnose Rays. PLoS ONE, 7 (7): e41605

YATES, P.M. & HEUPEL, M.R. & TOBIN, A.J. & SIMPFENDORFER, C.A. 2012 Diversity in young shark habitats provides the potential for portfolio effects. Marine Ecology Progress Series, 458: 269-281

YIGIN, C.C. & ISMEN, A. 2012 Age, growth and reproduction of the common stingray, Dasyatis pastinaca from the North Aegean Sea. Marine Biology Research, 8 (7): 644-653







 CABRERA, D.A. & CIONE, A.L. & BARLA, M.J. 2012 Three dimensional angel shark jaw elements (Elasmobranchii, Squatinidae) from the Miocene of Southern Argentina. Ameghiniana, 49 (1): 113-118

CIONE, A.L. & CABRERA, D.A. & BARLA, M.J. 2012 Oldest record of the Great White Shark (Lamnidae, Carcharodon; Miocene) in the Southern Atlantic. Geobios, 45 (2): 167-172

DUFFIN, C.J. 2012 Natternzungen-Kredenz: tableware for the Renaissance nobility. Jewellery History Today, 14: 3-5

HERMAN, J. & VAN WAES, H. 2012 Observations concernant l’Evolution et la Systématique de quelques Euselachii, Neoselachii et Batoidei (Pisces - Elasmobranchii), actuels et fossiles. Géominpal Belgica 2

HERMAN, J. & VAN WAES, H. 2012 Observations diverses et déductions concernant l’Evolution et la Systématique de quelques Euselachii, Neoselachii et Batoidei (Pisces - Elasmobranchii), actuels et fossiles. (Données complémentaires). Géominpal Belgica 2, 13pp

HERMAN, J. & VAN WAES, H. 2012 Observations diverses et déductions concernant l’Evolution et la Systématique de quelques Euselachii, Neoselachii et Batoidei (Pisces - Elasmobranchii), actuels et fossiles. (Fin) Géominpal Belgica 2

HODNETT, J.-P. & ELLIOTT, D.K. & OLSON, T.J. & WITTKE, J.H. 2012 Ctenacanthiform sharks from the Permian Kaibab Formation, northern Arizona. Historical Biology, 24 (4): 1-15

KRIWET, J. & KLUG, S. 2012 Presence of the extinct sawfish, Onchosaurus (Neoselachii, Sclerorhynchiformes) in the Late Cretaceous of Peru with a review of the genus. Journal of South American Earth Sciences, in press

VARONE, G. 2012 Note sur le genre Gymnura (Van Hasselt, 1823) [Poissons Chondrichthyens] présent à l’Oligocène dans le Bassin aquitain. Bulletin de la Société Linnéenne de Bordeaux, 40 (1): 81-83