Material: male, 48 cm TL, holotype, Bengale, Indian Ocean
Images: © Jacques Herman, Beigem (Grimbergen), Belgique, images shows the correct position of the teeth
Description after Springer, 1964 [#2274]: teeth smooth-edged, upper teeth 12-1-12 to 16-1-16; lower teeth 12-12 to 17-17 (higher, or lower, counts in upper and lower jaws correlated) ; enlarged hyomandibular pores 0-5 on each side of head (rarely more than 2; frequently difficult to see). Denticles imbricate, 3-ridged and 3-toothed in young, 3- to 5-ridged and 3- to 4-toothed in adults.
Sexual dimorphism: There is a striking difference in dentition of adult males as compared with females or juvenile males (fig. 3). In adult males the cusps of the teeth are long and slender and the middle and posterior teeth of each side of the lower jaw are erect. The anterior teeth are greatly elongate and flexuous and somewhat round in cross section. Several rows of the anterior lower teeth in the adult male appear to be functional.