Iglésias, S.P. & Lecointre, G. & Sellos, D.Y. (2005)
Extensive paraphylies within sharks of the order Carcharhiniformes inferred from nuclear and mitochondrial genes.
Molecular Phylogenetics and Evolution, 34(3), 569–583 DOI: 10.1016/j.ympev.2004.10.022