Furiness, S. & Williams, J.I. & Nagasawa, K. & Burreson, E.M. (2007)
A collection of fish leeches (Hirudinida: Piscicolidae) from Japan and surrounding waters, including redescriptions of three species.
Journal of Parasitology, 93(4), 875–883 DOI: 10.1645/GE-979R1.1