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Internal organs and body tissues of free-swimming whale sharks (Rhincodon typus) imaged using underwater ultrasound. Frontiers in Marine Science, 11, Article 1285429
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Asymptotic Growth of Whale Sharks Suggests Sex-Specific Life-History Strategies. Frontiers in Marine Science, 7, Article 575683
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Error and bias in size estimates of whale sharks: implications for understanding demography. Royal Society Open Science, 3(3), Article 150668
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The development of robust morphometric indices from accurate and precise measurements of free-swimming whale sharks using laser photogrammetry. Journal of the Marine Biological Association of the United Kingdom, 93(2), 309–320
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Long-term membership of whale sharks (Rhincodon typus) in coastal aggregations in Seychelles and Djibouti. Marine and Freshwater Research, 62(6), 621–627
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Seeing Spots: Photo-identification as a Regional Tool for Whale Shark Identification. Western Indian Ocean Journal of Marine Science, 9(2), 185–194