Vázquez-Liñero, M.A. & Galván-Magaña, F. & Tripp-Valdez, A. & Sánchez-González, A. & Briones-Hernández, S.A. & Hoyos-Padilla, E.M. & Piñón-Gimate, A. (2025)
Trophic ecology of juvenile smooth hammerhead shark Sphyrna zygaena (Carcharhiniformes: Sphyrnidae) in the central Gulf of California, Mexico. Environmental Biology of Fishes, in press
DOI: 10.1007/s10641-024-01665-5

Lara-Lizardi, F. & Castro, E. & Barajas, V.L. & Morales, J.M. & Hoyos-Padilla, E.M. & Ketchum, J. (2024)
Seasonal occurrence and individual variability of bull sharks, Carcharhinus leucas, in a marine reserve of the southwestern Gulf of California. PeerJ, 12, Article e17192
DOI: 10.7717/peerj.17192

Rubin, R.D. & Kumli, K.R. & Klimley, A.P. & Stewart, J.D. & Ketchum, J.T. & Hoyos-Padilla, E.M. & Galván-Magaña, F. & Zavala-Jiménez, A.A. & Fonseca-Ponce, I.A. & Saunders, M. & Dominguez-Sanchez, P.S. & Ahuja, P. & Nevels, C.R. & González, P.A.P. & Corgos, A. & Diemer, S.J. (2024)
Insular and mainland interconnectivity in the movements of oceanic manta rays (Mobula birostris) off Mexico in the Eastern Tropical Pacific. Environmental Biology of Fishes, in press
DOI: 10.1007/s10641-024-01622-2

Cabral, M.M.P. & Stewart, J.D. & Marques, T.A. & Ketchum, J.T. & Ayala-Bocos, A. & Hoyos-Padilla, E.M. & Reyes-Bonilla, H. (2023)
The influence of El Nino Southern Oscillation on the population dynamics of oceanic manta rays in the Mexican Pacific. Hydrobiologia, 850, 257–267
DOI: 10.1007/s10750-022-05047-9

Hoyos-Padilla, E.M. & Casanova-Santamaria, I. & Loria-Correa, J.C. & Sulikowski, J. (2023)
The successful use of a submersible ultrasound to confirm pregnancy on free swimming bull sharks, Carcharhinus leucas, in a provisioned shark site. Frontiers in Marine Science, 10, Article 1193563
DOI: 10.3389/fmars.2023.1193563

Santana-Morales, O. & Zertuche-Chanes, R. & Hoyos-Padilla, E.M. & Sepulveda, C. & Becerril-Garcia, E.E. & Gallo-Reynoso, J.P. & Barba-Acuna, I. & Mejia-Trejo, A. & Aquino-Baleyto, M. & Sosa-Nishizaki, O. & Ketchum, J.T. & Beas-Luna, R. (2022)
An exploration of the population characteristics and behaviours of the white shark in Guadalupe Island, Mexico (2014-2019): Observational data from cage diving vessels. Aquatic Conservation, Marine and Freshwater Ecosystems, 31(12), 3480–3491
DOI: 10.1002/aqc.3734

Klimley, A.P. & Ketchum, J.T. & Lara-Lizardi, F. & Papastamatiou, Y.P. & Hoyos-Padilla, E.M. (2022)
Evidence for spatial and temporal resource partitioning of sharks at Roca Partida, an isolated pinnacle in the eastern Pacific. Environmental Biology of Fishes, 105, 1963–1974
DOI: 10.1007/s10641-021-01206-4

Becerril-García, E.E. & Arauz, R. & Arellano-Martínez, M. & Bonfil, R. & Ayala-Bocos, A. & Castillo-Géniz, J.L. & Carrera-Fernández, M. & Charvet, P. & Chiaramonte, G. & Cisneros-Montemayor, A.M. & Concha, F. & Espinoza, M. & Ehemann, N.R. & Estupiñán-Montaño, C. & Fuentes, K. & Galván-Magaña, F. & Graham, R. & Hacohen-Domené, A. & Hazin, F. & Hernández, S. & Hoyos-Padilla, E.M. & Ketchum, J.T. & Kingma, I. & Méndez, O. & Oddone, M.C. & Pérez-Jiménez, J.C. & Petatán-Ramírez, D. & Polo-Silva, C. & Rangel, B. & Salinas-De-León, P. & Santana-Morales, O. & Zanella, I. & Vélez-Zuazo, X. & Godard-Codding, C.A.G. (2022)
Research priorities for the conservation of chondrichthyans in Latin America. Biological Conservation, 269, Artivle 109535
DOI: 10.1016/j.biocon.2022.109535

Lara-Lizardi, F. & Hoyos-Padilla, E.M. & Klimley, A.P. & Grau, M. & Ketchum, J.T. (2022)
Movement patterns and residency of bull sharks, Carcharhinus leucas, in a marine protected area of the Gulf of California. Environmental Biology of Fishes, 105, 1765–1779
DOI: 10.1007/s10641-022-01223-x

Malpica-Cruz, L. & Abadia-Cardoso, A. & Aquino-Baleyto, M. & Beas-Luna, R. & Becerril-Garcia, E.E. & Castillo-Geniz, J.L. & Galvan-Magana, F. & Garcia-Rodriguez, E. & Herzka, S.Z. & Hoyos-Padilla, E.M. & Lara-Mendoza, R.E. & Lorda, J. & Onate-Gonzalez, E.C. & Perez-Weil, R.J. & Saavedra-Sotelo, N.C. & Santana-Morales, O. & Towns, V. & Zepeda-Dominguez, J.A. (2021)
Empowering fishers for Great White Shark stewardship: Reply to Madigan et al. 2021. Conservation Letters, 14(4), Article e12828
DOI: 10.1111/conl.12828

Palacios, M.D. & Hoyos-Padilla, E.M. & Trejo-Ramírez, A. & Croll, D.A. & Galván-Magaña, F. & Zilliacus, K.M. & O'Sullivan, J.B. & Ketchum, J.T. & González-Armas, R. (2021)
Description of first nursery area for a pygmy devil ray species (Mobula munkiana) in the Gulf of California, Mexico. Scientific Reports, 11, Article 132
DOI: 10.1038/s41598-020-80506-8 PMCID: PMC7794486

Ayres, K.A. & Ketchum, J.T. & Gonzalez-Armas, R. & Galvan-Magana, F. & Hearn, A. & Elorriaga-Verplancken, F.R. & Hoyos-Padilla, E.M. & Kajiura, S.M. (2021)
The use of an unoccupied aerial vehicle to survey shark species over sand and rocky-reef habitats in a marine protected area. Journal of Fish Biology, 99(5), 1735–1740
DOI: 10.1111/jfb.14838

Santana-Morales, O. & Hoyos-Padilla, E.M. & Medellin-Ortiz, A. & Sepulveda, C. & Beas-Luna, R. & Aquino-Baleyto, M. & Becerril-Garcia, E.E. & Arellano-Millan, D. & Malpica-Cruz, L. & Lorda, J. & Castillo-Geniz, J.L. (2021)
How much is too much? A carrying capacity study of white shark cage diving in Guadalupe Island, Mexico. Marine Policy, 131, Article 104588
DOI: 10.1016/j.marpol.2021.104588

Pelamatti, T. & Rios-Mendoza, L.M. & Hoyos-Padilla, E.M. & Galvan-Magana, F. & De Camillis, R. & Marmolejo-Rodriguez, A.J. & Gonzalez-Armas, R. (2021)
Contamination knows no borders: Toxic organic compounds pollute plastics in the biodiversity hotspot of Revillagigedo Archipelago National Park, Mexico. Marine Pollution Bulletin, 170, Article 112623
DOI: 10.1016/j.marpolbul.2021.112623

Bessudo, S. & Ladino, F. & Becerril-Garcia, E.E. & Shepard, C.M. & Salinas-De-Leon, P. & Hoyos-Padilla, E.M. (2021)
The elasmobranchs of Malpelo Flora and Fauna Sanctuary, Colombia. Journal of Fish Biology, 99(5), 1769–1774
DOI: 10.1111/jfb.14874

Ayres, K.A. & Ketchum, J.T. & González-Armas, R. & Galván-Magaña, F. & Hearn, A. & Elorriaga-Verplancken, F.R. & Martínez-Rincón, R.O. & Hoyos-Padilla, E.M. & Kajiura, S.M. (2021)
Seasonal aggregations of blacktip sharks Carcharhinus limbatus at a marine protected area in the Gulf of California, assessed by unoccupied aerial vehicle surveys. Marine Ecology Progress Series, 678, 95–107
DOI: 10.3354/meps13897

Becerril-García, E.E. & Hoyos-Padilla, E.M. & Henning, B. & Salinas-de León, P. (2020)
Sharks, rays, and chimaeras of the Revillagigedo National Park: An update of new and confirmed records. Journal of Fish Biology, 97(4), 1228–1232
DOI: 10.1111/jfb.14457

Becerril-García, E.E. & Hoyos-Padilla, E.M. & Micarelli, P. & Galván-Magaña, F. & Sperone, E. (2020)
Behavioural responses of white sharks to specific baits during cage diving ecotourism. Scientific Reports, 10, Article 11152
DOI: 10.1038/s41598-020-67947-x

Becerril-García, E.E. & Arellano-Martínez, M. & Bernot-Simon, D. & Hoyos-Padilla, E.M. & Galván-Magaña, F. & Godard-Codding, C. (2020)
Steroid hormones and chondrichthyan reproduction: physiological functions, scientific research, and implications for conservation. PeerJ, 8, Article e9686
DOI: 10.7717/peerj.9686

Alderete-Macal, M.J. & Caraveo-Patino, J. & Hoyos-Padilla, E.M. (2020)
Ontogenetic differences in muscle fatty acid profile of white sharks Carcharodon carcharias off Guadalupe Island, Mexico. Revista de Biología Marina y Oceanografía, 55(1), 37–46
DOI: 10.22370/rbmo.2020.55.1.2372

Becerril-Garcia, E.E. & Hoyos-Padilla, E.M. & Santana-Morales, O. & Gutierrez-Ortiz, M.A. & Ayala-Bocos, A. & Galvan-Magana, F. (2020)
An estimate of the number of white sharks Carcharodon carcharias interacting with ecotourism in Guadalupe Island. Journal of Fish Biology, 97(6), 1861–1864
DOI: 10.1111/jfb.14540

Becerril-García, E.E. & Bernot-Simon, D. & Arellano-Martínez, M. & Galván-Magaña, F. & Santana-Morales, O. & Hoyos-Padilla, E.M. (2020)
Evidence of interactions between white sharks and large squids in Guadalupe Island, Mexico. Scientific Reports, 10, Article 17158
DOI: 10.1038/s41598-020-74294-4

Becerril-García, E.E. & Martínez-Rincón, R.O. & Galván-Magaña, F. & Santana-Morales, O. & Hoyos-Padilla, E.M. (2020)
Statistical modelling reveals spatial, temporal, and environmental preferences for white sharks at an oceanic aggregation site. Marine Ecology Progress Series, 655, 171–183
DOI: 10.3354/meps13528

Le Croizier, G. & Lorrain, A. & Sonke, J.E. & Hoyos-Padilla, E.M. & Galván-Magaña, F. & Santana-Morales, O. & Aquino-Baleytó, M. & Becerril-García, E.E. & Muntaner-López, G. & Ketchum, J. & Block, B. & Carlisle, A. & Jorgensen, Sj. & Besnard, L. & Jung, A. & Schaal, G. & Point, D. (2020)
The Twilight Zone as a Major Foraging Habitat and Mercury Source for the Great White Shark. Environmental Science & Technology, 54(24), 15872–15882
DOI: 10.1021/acs.est.0c05621

Becerril-Garcia, E.E. & Hoyos-Padilla, E.M. & Petatan-Ramirez, D. & Galvan-Magana, F. (2019)
Southernmost record of the white shark Carcharodon carcharias (Chondrichthyes: Lamnidae) in the Mexican Pacific. Latin American Journal of Aquatic Research, 47(1), 190–193
DOI: 10.3856/vol47-issue1-fulltext-22

Becerril-García, E.E. & Hoyos-Padilla, E.M. & Micarelli, P. & Galván-Magaña, F. & Sperone, E. (2019)
The surface behaviour of white sharks during ecotourism: A baseline for monitoring this threatened species around Guadalupe Island, Mexico. Aquatic Conservation, Marine and Freshwater Ecosystems, 29(5), 773–782
DOI: 10.1002/aqc.3057

Pazmiño, D.A. & Maes, G.E. & Green, M.E. & Simpfendorfer, C.A. & Hoyos-Padilla, E.M. & Duffy, C.J.A. & Meyer, C.G. & Kerwath, S.E. & Salinas-De-León, P. & Van Herwerden, L. (2018)
Strong trans-Pacific break and local conservation units in the Galapagos shark (Carcharhinus galapagensis) revealed by genome-wide cytonuclear markers. Journal of Heredity, 120(5), 407–421
DOI: 10.1038/s41437-017-0025-2

Arnes-Urgelles, C. & Hoyos-Padilla, E.M. & Pochet, F. & Salinas-De-Leon, P. (2018)
First observation on the mating behaviour of the marbled ray, Taeniurops meyeni, in the tropical Eastern Pacific. Environmental Biology of Fishes, 101(12), 1693–1699
DOI: 10.1007/s10641-018-0818-z

Huveneers, C. & Apps, K. & Becerril-García, E.E. & Bruce, B. & Butcher, P.A. & Carlisle, A.B. & Chapple, T.K. & Christiansen, H.M. & Cliff, G. & Curtis, T.H. & Daly-Engel, T.S. & Dewar, H. & Dicken, M.L. & Domeier, M.L. & Duffy, C.A.J. & Ford, R. & Francis, M.P. & French, G.C.A. & Galván-Magaña, F. & García-Rodríguez, E. & Gennari, E. & Graham, B. & Hayden, B. & Hoyos-Padilla, E.M. & Hussey, N.E. & Jewell, O.J.D. & Jorgensen, S.J. & Kock, A.A. & Lowe, C.G. & Lyons, K. & Meyer, L. & Oelofse, G. & Oñate-González, E.C. & Oosthuizen, H. & O'Sullivan, J.B. & Ramm, K. & Skomal, G. & Sloan, S. & Smale, M.J. & Sosa-Nishizaki, O. & Sperone, E. & Tamburin, E. & Towner, Av. & Wcisel, M.A. & Weng, K.C. & Werry, J.M. (2018)
Future Research Directions on the “Elusive” White Shark. Frontiers in Marine Science, 5, Article 455
DOI: 10.3389/fmars.2018.00455

Becerril-García, E.E & Hoyos-Padilla, E.M. & Micarelli, P. & Galván-Magaña, F. & Sperone, E. (2017)
Surf ace behavior of white shark, Carcharodon carcharias, related to legal and illegal baits in Guadalupe lsland, México [Abstract]. European Elasmobranch Association(EEA) 2017, Amsterdam

Salinas-De-Leon, P. & Hoyos-Padilla, E.M. & Pochet, F. (2017)
First observation on the mating behaviour of the endangered scalloped hammerhead shark Sphyrna lewini in the Tropical Eastern Pacific. Environmental Biology of Fishes, 100(12), 1603–1608
DOI: 10.1007/s10641-017-0668-0

Stewart, J.D. & Hoyos-Padilla, E.M. & Kumli, K.R. & Rubin, R.D. (2016)
Deep-water feeding and behavioral plasticity in Manta birostris revealed by archival tags and submersible observations. Zoology, 119(5), 406–413
DOI: 10.1016/j.zool.2016.05.010

Skomal, G.B. & Hoyos-Padilla, E.M. & Kukulya, A. & Stokey, R. (2015)
Subsurface observations of white shark Carcharodon carcharias predatory behaviour using an autonomous underwater vehicle. Journal of Fish Biology, 87(6), 1293–1312
DOI: 10.1111/jfb.12828

Sánchez-Hernández, X.E. & Díaz-Jaimes, P. & Uribe-Alcocer, M. & Hoyos-Padilla, E.M. & Blanco, P. (2014)
Estructura genética poblacional del tiburón blanco (Carcharodon carcharias) en el Pacifico Nororiental frente a las costas de Isla Guadalupe. In Libro de resúmenes, VI Simposium Nacional de Tiburones y Rayas, 07 al 12 de abril de 2014, Mazatlán, Sin: 65–67

Hoyos-Padilla, E.M. & Papastamatiou, Y.P. & O'Sullivan, J. & Lowe, C.G. (2013)
Observation of an Attack by a Cookiecutter Shark (Isistius brasiliensis) on a White Shark (Carcharodon carcharias). Pacific Science, 67(1), 129–134
DOI: 10.2984/67.1.10

Hoyos-Padilla, E.M. & Ceballos-Vázquez, B.P. & Galván-Magaña, F. (2012)
Reproductive Biology of the Silky Shark Carcharhinus falciformis (Chondrichthyes: Carcharhinidae) off the west coast of Baja California Sur, Mexico. Aqua, International Journal of Ichthyology, 18(1), 15–24

Galván-Magaña, F. & Hoyos-Padilla, E.M. & Navarro-Serment, C.J. & Márquez-Farías, F. (2010)
Records of white shark, Carcharodon carcharias, in the Gulf of California, Mexico. Marine Biodiversity Records, 3, Article e111
DOI: 10.1017/S1755267210000977

Hoyos-Padilla, E.M. (2003)
Biología reproductiva del tiburón piloto Carcharhinus falciformis (Bibron, 1839) de baja california sur. Thesis, Instituto Politécnico Nacional, Centro Interdisciplinario de Ciencias Marinas

Hoyos-Padilla, E.M. & Castillo-Géniz, J.L. (2000)
A review of the principal shark nurseries of the Gulf of México and Caribbean Sea [Abstract]. American Elasmobranch Society 16th Annual Meeting, 14–20 June 2000, La Paz, Mexico