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Only One Percent of Important Shark and Ray Areas in the Western Indian Ocean Are Fully Protected From Fishing Pressure. Ecology and Evolution, 16, Article 1
DOI: 10.1002/ece3.72690

Yu, C.J. & Joung, S.J. & Hsu, H.H. & Liu, K.M. & Yamaguchi, A. (2025)
Stable Isotope Analysis of Two Filter-Feeding Sharks in the Northwestern Pacific Ocean. Fishes, 10(6), Article 249
DOI: 10.3390/fishes10060249

Lin, Y.J. & Hsu, H.H. & Shiao, J.C. & Chung, M.T. & Rabaoui, L.J. (2025)
Specific locally, general globally: a synthesis on the isotopic niche in hammerhead sharks (Class Chondrichthyes, Genus Sphyrna). Reviews in Fish Biology and Fisheries, 35(3), 1609–1633
DOI: 10.1007/s11160-025-09974-3

Womersley, F.C. & Rohner, C.A. & Abrantes, K. & Afonso, P. & Arunrugstichai, S. & Bach, S.S. & Bar, S. & Barash, A. & Barnes, P. & Barnett, A. & Boldrocchi, G. & Buffat, N. & Canon, T. & Perez, C.C. & Chuangcharoendee, M. & Cochran, J.E.M. & de la Parra, R. & Diamant, S. & Driggers, W. & Dudgeon, C.L. & Erdmann, M. & Fitzpatrick, R. & Flam, A. & Fontes, J. & Francis, G. & Galvan, B.E. & Graham, R.T. & Green, S.M. & Green, J.R. & Grosmark, Y. & Guzman, H.M. & Hardenstine, R.S. & Harvey, M. & Harvey-Carroll, J. & Hasan, A.W. & Hearn, A.R. & Hendon, J.M. & Putra, M.I.H. & Himawan, M.R. & Hoffmayer, E. & Holmberg, J. & Hsu, H.H. & Jaidah, M.Y. & Jansen, A. & Judd, C. & Kuguru, B. & Lester, E. & Macena, B.C.L. & Magson, K. & Maguiño, R. & Manjaji-Matsumoto, M. & Marcoux, S.D. & Marcoux, T. & McKinney, J. & Meekan, M. & Mendoza, A. & Moazzam, M. & Monacella, E. & Norman, B. & Perry, C. & Pierce, S. & Prebble, C. & Macías, D.R. & Raudino, H. & Reynolds, S. & Robinson, D. & Rowat, D. & Santos, M.D. & Schmidt, J. & Scott, C. & See, S.T. & Sianipar, A. & Speed, C.W. & Syakurachman, I. & Tyne, J.A. & Waples, K. & Winn, C. & Yuneni, R.R. & Zareer, I. & Araujo, G. (2024)
Identifying priority sites for whale shark ship collision management globally. Science of the Total Environment, 934, Article 172776
DOI: 10.1016/j.scitotenv.2024.172776

Lin, Y.J. & Hsu, H.H. & Nazeer, Z. & Panickan, P. & Maneja, R.H. & Lozano-Cortés, D. & Qasem, A. & Huertas, A.D. & Rabaoui, L.J. (2024)
Resource use by Sphyrna mokarran and S. lewini (Chondrichthyes) neonates and juveniles in the western Arabian Gulf: a stable isotope analysis. Endangered Species Research, 54, 277–284
DOI: 10.3354/esr01341

Hsu, H.H. & Almealla, R. (2024)
7 | Regional Overviews, Indian Ocean, Bahrain In R.W. Jabado, A.Z.A. Morata, R.H. Bennett, B. Finucci, J.R. Ellis, S. Fowler, M.I. Grant &. A.P. Barbosa Martins, S.L. Sinclair (Eds.), The global status of sharks, rays, and chimaeras (pp. 1410–1416). International Union for Conservation of Nature and Natural Resources.
DOI: 10.59216/ssg.gsrsrc.2024

Hsu, H.H. (2024)
7 | Regional Overviews, Indian Ocean, Saudi Arabia, ARABIAN GULF In R.W. Jabado, A.Z.A. Morata, R.H. Bennett, B. Finucci, J.R. Ellis, S. Fowler, M.I. Grant &. A.P. Barbosa Martins, S.L. Sinclair (Eds.), The global status of sharks, rays, and chimaeras (pp. 1549–1554). International Union for Conservation of Nature and Natural Resources.
DOI: 10.59216/ssg.gsrsrc.2024

Hsu, H.H. & Tsai, W.-P. & Liu, K.-M. (2024)
7 | Regional Overviews, Indian Ocean, Taiwan, Province of China In R.W. Jabado, A.Z.A. Morata, R.H. Bennett, B. Finucci, J.R. Ellis, S. Fowler, M.I. Grant &. A.P. Barbosa Martins, S.L. Sinclair (Eds.), The global status of sharks, rays, and chimaeras (pp. 1769–1778). International Union for Conservation of Nature and Natural Resources.
DOI: 10.59216/ssg.gsrsrc.2024

Yacoubi, L. & El Zrelli, R.B. & Hsu, H.H. & Lin, Y.J. & Savoca, D. & Gopalan, J. & Nazal, M. & Bhuyan, M.S. & Arculeo, M. & Rabaoui, L.J. (2023)
Bioaccumulation of trace elements and hydrocarbons in chondrichthyans of the western Arabian Gulf: Environmental and human health risk assessment and implications for conservation. Science of the Total Environment, 901, Article 165990
DOI: 10.1016/j.scitotenv.2023.165990

Alhajji, A.H. & Hsu, H.H. & Alkhamis, Y.A. & Alsaqufi, A.S. & Khan, S.A. & Nazeer, Z. (2022)
Maturity and reproduction in the Arabian carpet shark, Chiloscyllium arabicum from the Saudi Arabian waters of the Arabian Gulf. Marine Biology Research, 18(5-6), 361–371
DOI: 10.1080/17451000.2022.2131824

Hsu, H.H. & Yacoubi, L. & Lin, Y.J. & Le Loc'h, F. & Katsanevakis, S. & Giovos, I. & Qurban, M.A. & Nazeer, Z. & Panickan, P. & Maneja, R.H. & Prihartato, P.K. & Loughland, R.A. & Rabaoui, L.J. (2022)
Elasmobranchs of the western Arabian Gulf: Diversity, status, and implications for conservation. Regional Studies in Marine Science, 56, Article 102637
DOI: 10.1016/j.rsma.2022.102637

Hsu, H.H. & Nazeer, Z. & Panickan, P. & Lin, Y.J. & Qasem, A. & Rabaoui, L.J. & Qurban, M.A. (2022)
Stomach Content Analysis for Juvenile Great Hammerhead Sharks Sphyrna mokarran (Ruppell, 1837) from the Arabian Gulf. Fishes, 7(6), Article 359
DOI: 10.3390/fishes7060359

Liu, K.M. & Huang, Y.W. & Hsu, H.H. (2021)
Management Implications for Skates and Rays Based on Analysis of Life History Parameters. Frontiers in Marine Science, 8, Article 664611
DOI: 10.3389/fmars.2021.664611

Yu, C.J. & Joung, S.J. & Hsu, H.H. & Lin, C.Y. & Hsieh, T.C. & Liu, K.M. & Yamaguchi, A. (2021)
Spatial-Temporal Distribution of Megamouth Shark, Megachasma pelagios, Inferred from over 250 Individuals Recorded in the Three Oceans. Animals, 11, Article 2947
DOI: 10.3390/ani11102947

Ong, J.J.L. & Meekan, M.G. & Hsu, H.H. & Fanning, L.P. & Campana, S.E. (2020)
Annual Bands in Vertebrae Validated by Bomb Radiocarbon Assays Provide Estimates of Age and Growth of Whale Sharks. Frontiers in Marine Science, 7, Article 188
DOI: 10.3389/fmars.2020.00188

Hsu, H.H. & Nazeer, Z.M. & Lin, Y.J. & Panickan, P. & Al-Abdulkader, K. & Loughland, R. & Qurban, M.A. (2020)
Biological aspects of juvenile great hammerhead sharks Sphyrna mokarran from the Arabian Gulf. Marine and Freshwater Research, 72(1), 110–117
DOI: 10.1071/MF19368

Joung, S.J. & Lyu, G.T. & Su, K.Y. & Hsu, H.H. & Liu, K.M. (2017)
Distribution Pattern, Age, and Growth of Blue Sharks in the South Atlantic Ocean. Marine and Coastal Fisheries, 9(1), 38–49
DOI: 10.1080/19425120.2016.1249579

Chang, C.-H. & Jang-Liaw, N.H. & Lin, Y.S. & Carlisle, A.B. & Hsu, H.H. & Liao, Y.C. & Shao, K.T. (2016)
The complete mitochondrial genome of the salmon shark, Lamna ditropis (Chondrichthyes, Lamnidae). Mitochondrial DNA, 27(1), 316–317
DOI: 10.3109/19401736.2014.892095

Hsu, H.H. & Lin, C.Y. & Joung, S.J. (2014)
The first record, tagging and release of a neonatal whale shark Rhincodon typus in Taiwan. Journal of Fish Biology, 85(5), 1753–1756
DOI: 10.1111/jfb.12498

Hsu, H.H. & Joung, S.J. & Hueter, R.E. & Liu, K.-M. (2014)
Age and growth of the whale shark (Rhincodon typus) in the north-western Pacific. Marine and Freshwater Research, 65(12), 1145–1154
DOI: 10.1071/MF13330

Hsu, H.H. & Joung, S.J. & Ebert, D.A. & Lin, C.Y. (2013)
Records of new and rare elasmobranchs from Taiwan. Zootaxa, 3752, 249–255
DOI: 10.11646/zootaxa.3752.1.15

Hsu, H.H. & Leu, M.-Y. & Wang, W.-H. & Hsieh, T.-C. & Joung, S.J. (2013)
Tagging and Release of a Captive Whale Shark after Eight Years in Taiwan's Aquarium: A Case Study. Journal of the Fisheries Society of Taiwan, 40(3), 143–151

Hsu, H.H. & Joung, S.J. & Liu, K.-M. (2012)
Fisheries, management and conservation of the whale shark Rhincodon typus in Taiwan. Journal of Fish Biology, 80(5), 1595–1607
DOI: 10.1111/j.1095-8649.2012.03234.x

Hsu, H.H. & Joung, S.-J. & Liao, Y.-Y. & Liu, K.-M. (2007)
Satellite tracking of juvenile whale sharks, Rhincodon typus, in the Northwestern Pacific. Fisheries Research, 84(1), 25–31
DOI: 10.1016/j.fishres.2006.11.030

Joung, S.-J. & Liu, K.-M. & Liao, Y.-Y. & Hsu, H.H. (2005)
Observed by-catch of Taiwanese tuna longline fishery in the South Atlantic Ocean. Journal of the Fisheries Society of Taiwan, 32(1), 69–77

Joung, S.-J. & Hsu, H.H. (2005)
Reproduction and Embryonic Development of the Shortfin Mako, Isurus oxyrinchus Rafinesque, 1810, in the Northwestern Pacific. Zoological Studies, 44(4), 487–496

Joung, S.-J. & Liu, K.-M. & Hsu, H.H. (2004)
Observed by-catch of Taiwanese tuna longline fishery in Atlantic Ocean. ICCAT SCRS/2004/184. 12 pp.