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Assessing the potential of acoustic telemetry to underpin the regional management of basking sharks (Cetorhinus maximus). Animal Biotelemetry, 12(1), Article 20
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Klimley, A.P. & Curtis, T.H. & Johnston, E.M. & Kock, A. & Stevens, G.M.W. (2024)
A review of elasmobranch breaching behavior: why do sharks and rays propel themselves out of the water into the air? Environmental Biology of Fishes, in press
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Rubin, R.D. & Kumli, K.R. & Klimley, A.P. & Stewart, J.D. & Ketchum, J.T. & Hoyos-Padilla, E.M. & Galván-Magaña, F. & Zavala-Jiménez, A.A. & Fonseca-Ponce, I.A. & Saunders, M. & Dominguez-Sanchez, P.S. & Ahuja, P. & Nevels, C.R. & González, P.A.P. & Corgos, A. & Diemer, S.J. (2024)
Insular and mainland interconnectivity in the movements of oceanic manta rays (Mobula birostris) off Mexico in the Eastern Tropical Pacific. Environmental Biology of Fishes, in press
DOI: 10.1007/s10641-024-01622-2
Paez, A.M. & Padilla, E.M.H. & Klimley, A.P. (2023)
A review of tonic immobility as an adaptive behavior in sharks. Environmental Biology of Fishes, 106(6), 1455–1462
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Kanive, P.E. & Rotella, J.J. & Chapple, T.K. & Anderson, S.D. & Hoyos-Padilla, M. & Klimley, A.P. & Galvan-Magana, F. & Andrzejaczek, S. & Block, B.A. & Jorgensen, S.J. (2023)
Connectivity between white shark populations off Central California, USA and Guadalupe Island, Mexico. Frontiers in Marine Science, 10, Article 1210969
DOI: 10.3389/fmars.2023.1210969
Bravo-Ormaza, E. & Arauz, R. & Bessudo, S. & Hearn, A. & Klimley, A.P. & Ladino-Archila, F. & Lopez-Macias, J. & Steiner, T. & Penaherrera-Palma, C. (2023)
Scalloped hammerhead shark Sphyrna lewini relative abundance comparison in three offshore marine protected areas of the Eastern Tropical Pacific. Environmental Biology of Fishes, 106(8), 1767–1784
DOI: 10.1007/s10641-023-01454-6
Klimley, A.P. (2023)
A historical approach to describing the complex behaviour of a large species of carnivorous shark - Case study No. 1: the scalloped hammerhead, Sphyrna lewini. Behaviour, 160(11-14), 1081–1102
DOI: 10.1163/1568539X-bja10211
Klimley, A.P. (2023)
A historical approach to describing the complex behaviour of a large species of predatory shark - Case study 2: the white shark, Carcharodon carcharias. Behaviour, 160(11-14), 1103–1136
DOI: 10.1163/1568539X-bja10194
Klimley, A.P. & Hoyos-Padilla, M. (2023)
Could convulsive body shuddering of a white shark near a shark cage be an element of a threat display? Behaviour, 160(11-14), 1245–1257
DOI: 10.1163/1568539X-bja10201
Klimley, A.P. & Porcher, I.F. & Clua, E.E.G. & Pratt, H.L. & Jr. (2023)
A review of the behaviours of the Chondrichthyes: a multi-species ethogram for the chimaeras, sharks, and rays. Behaviour, 160(11-14), 967–1080
DOI: 10.1163/1568539X-bja10214
Klimley, A.P. & Arauz, R. & Bessudo, S. & Chavez, E.J. & Chinacalle, N. & Espinoza, E. & Green, J. & Hearn, A.R. & Hoyos-Padilla, M.E. & Nalesso, E. & Ketchum, J.T. & Penaherrera-Palma, C. (2022)
Studies of the movement ecology of sharks justify the existence and expansion of marine protected areas in the Eastern Pacific Ocean. Environmental Biology of Fishes, 105, 2133–2153
DOI: 10.1007/s10641-021-01204-6
Johnston, E.M. & Houghton, J.D.R. & Mayo, P.A. & Hatten, G.K.F. & Klimley, A.P. & Mensink, P.J. (2022)
Cool runnings: behavioural plasticity and the realised thermal niche of basking sharks. Environmental Biology of Fishes, 105, 2001–2015
DOI: 10.1007/s10641-021-01202-8
Klimley, A.P. & Ketchum, J.T. & Lara-Lizardi, F. & Papastamatiou, Y.P. & Hoyos-Padilla, E.M. (2022)
Evidence for spatial and temporal resource partitioning of sharks at Roca Partida, an isolated pinnacle in the eastern Pacific. Environmental Biology of Fishes, 105, 1963–1974
DOI: 10.1007/s10641-021-01206-4
Lara-Lizardi, F. & Hoyos-Padilla, E.M. & Klimley, A.P. & Grau, M. & Ketchum, J.T. (2022)
Movement patterns and residency of bull sharks, Carcharhinus leucas, in a marine protected area of the Gulf of California. Environmental Biology of Fishes, 105, 1765–1779
DOI: 10.1007/s10641-022-01223-x
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Reply to: Shark mortality cannot be assessed by fishery overlap alone. Nature, 595(7866), E8–E16
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Triennial migration and philopatry in the critically endangered soupfin shark Galeorhinus galeus. Journal of Applied Ecology, 58(8), 1570–1582
DOI: 10.1111/1365-2664.13848
Mcinturf, A.G. & Steel, A.E. & Buckhorn, M. & Sandstrom, P. & Slager, C.J. & Fangue, N.A. & Klimley, A.P. & Caillaud, D. (2019)
Use of a hydrodynamic model to examine behavioral response of broadnose sevengill sharks (Notorynchus cepedianus) to estuarine tidal flow. Environmental Biology of Fishes, 102(9), 1149–1159
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Global spatial risk assessment of sharks under the footprint of fisheries. Nature, 572(7770), 461
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Galván-Magaña, F. & Castillo-Geniz, J.L. & Hoyos-Padilla, M. & Ketchum, J. & Klimley, A.P. & Ramírez-Amaro, S. & Torres-Rojas, Y.E. & Tovar-Avila, J. (2019)
3. Shark ecology, the role of the apex predator and current conservation status. Advances in Marine Biology, 83, 61–114
DOI: 10.1016/bs.amb.2019.08.005
Gallagher, A.J. & Klimley, A.P. (2018)
The biology and conservation status of the large hammerhead shark complex: the great, scalloped, and smooth hammerheads. Reviews in Fish Biology and Fisheries, 28(4), 777–794
DOI: 10.1007/s11160-018-9530-5
Hoyos‑Padilla, E.M. & Klimley, A.P. & Galván-Magaña, F. & Antoniou, A. (2016)
Contrasts in the movements and habitat use of juvenile and adult white sharks (Carcharodon carcharias) at Guadalupe Island, Mexico. Animal Biotelemetry, 4, Article 14
DOI: 10.1186/s40317-016-0106-7
Hearn, A.R. & Green, J. & Roman, M.H. & Acuna-Marrero, D. & Espinoza, E. & Klimley, A.P. (2016)
Adult female whale sharks make long-distance movements past Darwin Island (Galapagos, Ecuador) in the Eastern Tropical Pacific. Marine Biology, 163(10), Article 214
DOI: 10.1007/s00227-016-2991-y
Klimley, A.P. (2015)
Shark Trails of the Eastern Pacific. American Scientist, 103(4), 276–283
Campana, S.E. & Fisk, A.T. & Klimley, A.P. (2015)
Movements of Arctic and northwest Atlantic Greenland sharks (Somniosus microcephalus) monitored with archival satellite pop-up tags suggest long-range migrations. Deep Sea Research Part II: Topical Studies in Oceanography, 115, 109–115
DOI: 10.1016/j.dsr2.2013.11.001
Kanive, P.E. & Rotella, J.J. & Jorgensen, S.J. & Chapple, T.K. & Anderson, S.D. & Klimley, A.P. & Block, B.A. (2015)
Estimating apparent survival of sub-adult and adult white sharks (Carcharodon carcharias) in central California using mark-recapture methods. Frontiers in Marine Science, 2, Article 19
DOI: 10.3389/fmars.2015.00019
Ketchum, J.T. & Hearn, A. & Klimley, A.P. & Espinoza, E. & Peñaherrera, C. & Largier, J.L. (2014)
Seasonal changes in movements and habitat preferences of the scalloped hammerhead shark (Sphyrna lewini) while refuging near an oceanic island. Marine Biology, 161(4), 755–767
DOI: 10.1007/s00227-013-2375-5
Ketchum, J.T. & Hearn, A. & Klimley, A.P. & Peñaherrera, C. & Espinoza, E. & Bessudo, S. & Soler, G. & Arauz, R. (2014)
Inter-island movements of scalloped hammerhead sharks (Sphyrna lewini) and seasonal connectivity in a marine protected area of the eastern tropical Pacific. Marine Biology, 161(4), 939–951
DOI: 10.1007/s00227-014-2393-y
Ketchum, J.T. & Hearn, A. & Klimley, A.P. & Espinoza, E. & Peñaherrera, C. & Bessudo, S. & Soler, G. & Arauz, R. & Chapple, T.K. & Hoyos-Padilla, M. (2014)
Shark movements and the design of pelagic marine reserves [Abstract]. In Programm and Abstracts of Shark International, Durban 2014: 109
Ketchum, J.T. & Galván-Magaña, F. & Klimley, A.P. (2013)
Segregation and foraging ecology of whale sharks, Rhincodon typus, in the southwestern Gulf of California. Environmental Biology of Fishes, 96(6), 779–795
DOI: 10.1007/s10641-012-0071-9
Anderson, S.D. & Chapple, T.K. & Jorgensen, S.J. & Klimley, A.P. & Block, B.A. (2011)
Long-term individual identification and site fidelity of white sharks, Carcharodon carcharias, off California using dorsal fins. Marine Biology, 158(6), 1233–1237
DOI: 10.1007/s00227-011-1643-5
Chapple, T.K. & Jorgensen, S.J. & Anderson, S.D. & Kanive, P.E. & Klimley, A.P. & Botsford, L.W. & Block, B.A. (2011)
A first estimate of white shark, Carcharodon carcharias, abundance off Central California. Biology Letters, 7(4), 581–583
DOI: 10.1098/rsbl.2011.0124
Bessudo, S. & Soler, G.A. & Klimley, A.P. & Ketchum, J.T. & Hearn, A. & Arauz, R. (2011)
Residency of the scalloped hammerhead shark (Sphyrna lewini) at Malpelo Island and evidence of migration to other islands in the Eastern Tropical Pacific. Environmental Biology of Fishes, 91(2), 165–176
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Jorgensen, S.J. & Reeb, C.A. & Chapple, T.K. & Anderson, S.D. & Perle, C. & Van Sommeran, S.R. & Fritz-Cope, C. & Brown, A.C. & Klimley, A.P. & Block, B.A. (2010)
Philopatry and migration of Pacific white sharks. Proceedings of the Royal Society of London, Series B, 277, 1–10
DOI: 10.1098/rspb.2009.1155
Hearn, A. & Ketchum, J. & Klimley, A.P. & Espinoza, E. & Peñaherrera, C. (2010)
Hotspots within hotspots? Hammerhead shark movements around Wolf Island, Galapagos Marine Reserve Marine Biology, 157(9), 1899–1915
DOI: 10.1007/s00227-010-1460-2
Jorgensen, S.J. & Klimley, A.P. & Muhlia-Melo, A.F. (2009)
Scalloped hammerhead shark Sphyrna lewini, utilizes deep-water, hypoxic zone in the Gulf of California. Journal of Fish Biology, 74(7), 1682–1687
DOI: 10.1111/j.1095-8649.2009.02230.x
Klimley, A.P. & Curtis, T.H. (2007)
Shark Attack versus Ecotourism: Negative and Positive Interactions. In TIMM, R. M. & O'BRIEN, J. M., eds. Proceedings of the 22nd Vertebrate Pest Conference, 2006, University of California: 33–44
Curtis, T.H. & Kelly, J.T. & Menard, K.L. & Laroche, R.K. & Jones, R.E. & Klimley, A.P. (2006)
Observations on the Behavior of White Sharks Scavenging from a Whale Carcass at Point Reyes, California. California Fish and Game, 92(3), 113–124
Klimley, A.P. & Kihslinger, R.L. & Kelly, J.T. (2005)
Directional and Non-directional Movements of Bat Rays, Myliobatis californica, in Tomales Bay, California. Environmental Biology of Fishes, 74(1), 79–88
DOI: 10.1007/s10641-005-3997-3
Klimley, A.P. & Beavers, S.C. & Curtis, T.H. & Jorgensen, S.J. (2002)
Movements and Swimming Behavior of Three Species of Sharks in La Jolla Canyon, California. Environmental Biology of Fishes, 63(2), 117–135
DOI: 10.1023/A:1014200301213
Klimley, A.P. & Le Boeuf, B.J. & Cantara, K.M. & Richert, J.E. & Davis, S.F. & Sommeran, S.V. (2001)
Radio-acoustic positioning as a tool for studying site-specific behavior of the white shark and other large marine species. Marine Biology, 138(2), 429–446
DOI: 10.1007/s002270000394
Klimley, A.P. & Le Boeuf, B.J. & Cantara, K.M. & Richert, J.E. & Davis, S.F. & Sommeran, S.V. & Kelly, J.T. (2001)
The hunting strategy of white sharks (Carcharodon carcharias) near a seal colony. Marine Biology, 138(3), 617–636
DOI: 10.1007/s002270000489
Goldman, K.J. & McCosker, J.E. & Anderson, S.D. & Klimley, A.P. (1996)
Temperature, Swimming Depth, and Movements of a White Shark at the South Farallon Islands, California. In A.P. Klimley & D. Ainley, (Eds.), Great White Sharks. The biology of Carcharodon carcharias (pp. 111–120). Academic Press.
Klimley, A.P. & Ainley, D.G. (1996)
Great White Sharks. Academic Press. 517pp
Mollet, H.F. & Ebert, D.A. & Cailliet, G.M. & Testi, A.D. & Klimley, A.P. & Compagno, L.J.V. (1996)
A review of length validation methods and protocols to measure large white sharks. In A.P. Klimley & D. Ainley, (Eds.), Great White Sharks. The biology of Carcharodon carcharias (pp. 91–108). Academic Press.
Klimley, A.P. & Pyle, P. & Anderson, S.D. (1996)
Tail Slap and Breach Agonistic Displays among White Sharks? In A.P. Klimley & D. Ainley, (Eds.), Great White Sharks. The biology of Carcharodon carcharias (pp. 241–255). Academic Press.
Anderson, S.D. & Klimley, A.P. & Pyle, P. & Henderson, R.P. (1996)
Tidal Height and White Shark Predation at the Farallon Islands, California. In A.P. Klimley & D. Ainley, (Eds.), Great White Sharks. The biology of Carcharodon carcharias (pp. 275–279). Academic Press.
Klimley, A.P. & Ainley, D.G. (1996)
White Shark Research in the Past A Perspective. In A.P. Klimley & D. Ainley, (Eds.), Great White Sharks. The biology of Carcharodon carcharias (pp. 3–4). Academic Press.
Sillman, A.J. & Letsinger, G.A. & Patel, S. & Loew, E.R. & Klimley, A.P. (1996)
Visual pigments and photoreceptors in two species of shark, Triakis semifasciata and Mustelus henlei. Journal of Experimental Zoology, 276(1), 1–10
Klimley, A.P. & Pyle, P. & Anderson, S.D. (1996)
The Behavior of White Sharks and Their Pinniped Prey during Predatory Attacks. In A.P. Klimley & D. Ainley, (Eds.), Great White Sharks. The biology of Carcharodon carcharias (pp. 175–191). Academic Press.
Pyle, P. & Klimley, A.P. & Anderson, S.D. & Henderson, R.P. (1996)
Environmental Factors Affecting the Occurrence and Behavior of White Sharks at the Farallon Islands, California. In A.P. Klimley & D. Ainley, (Eds.), Great White Sharks. The biology of Carcharodon carcharias (pp. 281–291). Academic Press.
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Residency Patterns of White Sharks at the South Farallon Islands, California. In A.P. Klimley & D. Ainley, (Eds.), Great White Sharks. The biology of Carcharodon carcharias (pp. 365–373). Academic Press.
Klimley, A.P. (1994)
The predatory behavior of the white shark. American Scientist, 82(2), 122–133
Klimley, A.P. (1994)
Do white sharks (Carcharodon carcharias) select prey based upon high fat content? Abstract American Elasmobranch Society 10th Annual Meeting, 2.–8. June 1994, University of Southern California, Los Angeles
Mollet, H.F. & Cailliet, G.M. & Klimley, A.P. & Ebert, D.A. & Testi, A.D. (1994)
Length validation of the Kangaroo Island and Malta white sharks, Carcharodon carcharias, with estimated total length of 7 m [Abstract]. American Elasmobranch Society 10th Annual Meeting, 2.–8. June 1994, University of Southern California, Los Angeles
Klimley, A.P. & Cabrera-Mancilla, I. & Castillo-Géniz, J.L. (1993)
Horizontal and vertical movements of the Scalloped Hammerhead Shark, Sphyrna lewini, in the Southern Gulf of California, Mexico. Ciencias Marinas, 19(1), 95–115
Klimley, A.P. (1993)
Highly directional swimming by scalloped hammerhead sharks, Sphyrna lewini, and subsurface irradiance, temperature, bathymetry, and geomagnetic field. Marine Biology, 117(1), 1–22
DOI: 10.1007/BF00346421
Klimley, A.P. & Anderson, S.D. & Pyle, P. & Henderson, R.P. (1992)
Spatiotemporal patterns of white shark (Carcharodon carcharias) predation at the South Farallon Islands, California. Copeia, 1992(3), 680–690
Galván-Magaña, F. & Nienhuis, H.J. & Klimley, A.P. (1989)
Seasonal abundance and feeding habits of sharks of the lower Gulf of California, Mexico. California Fish and Game, 75(2), 74–84
Klimley, A.P. & Butler, S.B. & Nelson, D.R. & Stull, A.T. (1988)
Diel movements of scalloped hammerhead sharks, Sphyrna lewini Griffith and Smith, to and from a seamount in the Gulf of California. Journal of Fish Biology, 33(5), 751–761
DOI: 10.1111/j.1095-8649.1988.tb05520.x
Klimley, A.P. & Butler, S.B. (1988)
Immigration and emigration of a pelagic fish assemblage to seamounts in the Gulf of California related to water mass movements using satellite imagery. Marine Ecology Progress Series, 49, 11–20
Klimley, A.P. (1987)
The determinants of sexual segregation in the scalloped hammerhead shark, Sphyrna lewini. Environmental Biology of Fishes, 18(1), 27–40
DOI: 10.1007/BF00002325
Klimley, A.P. (1985)
The areal distribution and autoecology of the white shark Carcharodon carcharias off the west coast of North America. Memoirs of the Southern California Academy of Sciences, 9, 15–40
Klimley, A.P. (1985)
Schooling in the large predator, Sphyrna lewini, a species with low risk of predation: a non-egalitarian state. Zeitschrift für Tierpsychology, 70(4), 297–319
DOI: 10.1111/j.1439-0310.1985.tb00520.x
Klimley, A.P. & Nelson, D.R. (1984)
Diel movement patterns of the scalloped hammerhead shark (Sphyrna lewini) in relation to El Bajo Espiritu Santo: a refuging central-position social system. Behavioral Ecology and Sociobiology, 15(1), 45–54
DOI: 10.1007/BF00310214
Klimley, A.P. (1983)
Social organization of schools of the scalloped hammerhead shark, Sphyrna lewini (Griffith and Smith), in the Gulf of California. Ph.D. Dissertation. University of California, San Diego. 339 p.
Klimley, A.P. & Brown, S.T. (1983)
Stereophotography for the field biologist: measurement of lengths and three-dimensional positions of free-swimming sharks. Marine Biology, 74(2), 175–185
DOI: 10.1007/BF00413921
Klimley, A.P. & Nelson, D.R. (1981)
Schooling of the scalloped hammerhead sharks, Sphyrna lewini, in the Gulf of California. Fishery Bulletin, 79(2), 356–360
Klimley, A.P. (1981)
Grouping behavior in the scalloped hammerhead. Oceanus, 24(4), 65–71
Klimley, A.P. (1980)
Observations of courtship and copulation in the nurse shark, Ginglymostoma cirratum. Copeia, 1980(4), 878–882
Klimley, A.P. & Myrberg, A.A. (1979)
Acoustic stimuli underlying withdrawal from a sound source by adult lemon sharks, Negaprion brevirostris (Poey). Bulletin of Marine Science, 29(4), 447–458
Klimley, A.P. (1978)
An inquiry into the causes of shark attack. Sea Frontiers, 20(2), 67–76
Klimley, A.P. (1976)
The white shark, a matter of size. Sea Frontiers, 22(1), 2–8
Klimley, A.P. (1974)
Nurses at home and school. Marine Aquarist, 8(6), 5–13