Homma, K. & Maruyama, T. & Itoh, T. & Ishihara, H. & Uchida, S. (1999)
Biology of the manta ray, Manta birostris Walbaum in the Indo-Pacific. In Seret, B. and Sire, J.Y. (Eds) Indo-Pacific fish biology: Proceedings of the 5th Indo-Pacific Fisheries Conference, Noumea, 1997. Ichthyological Society of France, Paris: 209–216
Homma, K. & Maruyama, T. & Takeda, Y. & Ishihara, H. (1994)
A study on the biology of rays occurring at Pohnpei Island, Caroline Islands. Proceedings of the 4th Indo-Pacific Fish Conference, Bankok: 87–107