Kanna, T. & Takahashi, S. & Byun, E. & Yamashiro, A. & Matsumoto, R. & Torisawa, S. & Mitsunaga, Y. (2025)
Seasonal behavioral changes of a captive whale shark (Rhincodon typus) under variable temperature and light conditions. Fishery Bulletin, 123(2), 75–87
DOI: 10.7755/fb.123.2.1

Meekan, M.G. & Thompson, F. & Brooks, K. & Matsumoto, R. & Murakumo, K. & Lester, E. & Dove, A. & Hopper, B. (2024)
Internal organs and body tissues of free-swimming whale sharks (Rhincodon typus) imaged using underwater ultrasound. Frontiers in Marine Science, 11, Article 1285429
DOI: 10.3389/fmars.2024.1285429

Inoue, T. & Shimoyama, K. & Saito, M. & Wong, M.K.S. & Ikeba, K. & Nozu, R. & Matsumoto, R. & Murakumo, K. & Sato, K. & Tokunaga, K. & Kofuji, K. & Takagi, W. & Hyodo, S. (2022)
Long-term monitoring of egg-laying cycle using ultrasonography reveals the reproductive dynamics of circulating sex steroids in an oviparous catshark, Scyliorhinus torazame. General and Comparative Endocrinology, 327, Article 114076
DOI: 10.1016/j.ygcen.2022.114076

Tomita, T. & Toda, M. & Murakumo, K. & Miyamoto, K. & Matsumoto, R. & Ueda, K. & Sato, K. (2021)
Volume of the whale shark and their mechanism of vertical feeding. Zoology, 147, Article 125932
DOI: 10.1016/j.zool.2021.125932

Yagishita, N. & Ikeguchi, S.-I. & Matsumoto, R. (2020)
Re-Estimation of Genetic Population Structure and Demographic History of the Whale Shark (Rhincodon typus) with Additional Japanese Samples, Inferred from Mitochondrial DNA Sequences. Pacific Science, 74(1), 31–47
DOI: 10.2984/74.1.3

Murakumo, K. & Matsumoto, R. & Tomita, T. & Matsumoto, Y. & Ueda, K. (2020)
The power of ultrasound: observation of nearly the entire gestation and embryonic developmental process of captive reef manta rays (Mobula alfredi). Fishery Bulletin, 118(1), 1–7
DOI: 10.7755/FB.118.1.1

Nakamura, I. & Matsumoto, R. & Sato, K. (2020)
Body temperature stability in the whale shark, the world's largest fish. Journal of Experimental Biology, 223(11), Article jeb210286
DOI: 10.1242/jeb.210286

Wyatt, A.S.J. & Matsumoto, R. & Chikaraishi, Y. & Miyairi, Y. & Yokoyama, Y. & Sato, K. & Ohkouchi, N. & Nagata, T. (2019)
Enhancing insights into foraging specialization in the world's largest fish using a multi-tissue, multi-isotope approach. Ecological Monographs, 89(1), Article e01339
DOI: 10.1002/ecm.1339

Matsumoto, R. & Matsumoto, Y. & Ueda, K. & Suzuki, M. & Asahina, K. & Sato, K. (2019)
Sexual maturation in a male whale shark (Rhincodon typus) based on observations made over 20 years of captivity. Fishery Bulletin, 117(1–2), 78–86
DOI: 10.7755/FB.117.1-2.9

Imaseki, I. & Wakabayashi, M. & Hara, Y. & Watanabe, T. & Takabe, S. & Kakumura, K. & Honda, Y. & Ueda, K. & Murakumo, K. & Matsumoto, R. & Matsumoto, Y. & Nakamura, M. & Takagi, W. & Kuraku, S. & Hyodo, S. (2019)
Comprehensive analysis of genes contributing to euryhalinity in the bull shark, Carcharhinus leucas; Na+-Cl- co-transporter is one of the key renal factors upregulated in acclimation to low-salinity environment. Journal of Experimental Biology, 222(12), Article jeb201780
DOI: 10.1242/jeb.201780

Nozu, R. & Matsumoto, R. & Murakumo, K. & Sato, K. (2019)
Observation in aquarium sheds light on reproductive biology oflarge elasmobranch fish [in Japanese]. Report of Japanese Society for Elasmobranch Studies, 55, 2–7

Nozu, R. & Murakumo, K. & Yano, N. & Furuyama, R. & Matsumoto, R. & Yanagisawa, M. & Sato, K. (2018)
Changes in sex steroid hormone levels reflect the reproductive status of captive female zebra sharks (Stegostoma fasciatum). General and Comparative Endocrinology, 265, 174–179
DOI: 10.1016/j.ygcen.2018.03.006

Hara, Y. & Yamaguchi, K. & Onimaru , K. & Kadota, M. & Koyanagi, M. & Keeley, S.D. & Tatsumi, K. & Tanaka, K. & Motone, F. & Kageyama, Y. & Nozu, R. & Adachi, N. & Nishimura, O. & Nakagawa, R. & Tanegashima, C. & Kiyatake, I. & Matsumoto, R. & Murakumo, K. & Nishida, K. & Terakita, A. & Kuratani, S. & Sato, K. & Hyodo, S. & Kuraku, S. (2018)
Shark genomes provide insights into elasmobranch evolution and the origin of vertebrates. Nature Ecology & Evolution, 2, 1761–1771

Matsumoto, R. & Toda, M. & Matsumoto, Y. & Ueda, K. & Nakazato, M. & Sato, K. & Uchida, S. (2017)
Chapter 2: Notes on Husbandry of Whale Sharks, Rhincodon typus, in Aquaria. In SMITH, M. & WARMOLTS, D. & THONEY, D. & HUETER, R. & MURRAY, M. & EZURRA, J.(EDITORS): 2017. The Elasmobranch Husbandry Manual II: Recent Advances in the Care of Sharks, Rays and their Relatives. Special Publication of the Ohio Biological Survey: 15-22

Nozu, R. & Murakumo, K. & Matsumoto, R. & Nakamura, M. & Ueda, K. & Sato, K. (2015)
Gonadal Morphology, Histology, and Endocrinological Characteristics of Immature Female Whale Sharks, Rhincodon typus. Zoological Science, 32(5), 455–458
DOI: 10.2108/zs150040

Nakaya, K. & Matsumoto, R. & Suda, K. (2008)
Feeding strategy of the megamouth shark Megachasma pelagios (Lamniformes: Megachasmidae). Journal of Fish Biology, 73(1), 17–34
DOI: 10.1111/j.1095-8649.2008.01880.x