The genus Heterodontus in the Early Paleogene of the North Sea Basin and the Landes, France. Palaeo Ichthyologica, 17, 1–60
On the dentition of the extant species Heterodontus mexicanus and Heterodontus quoyi. Palaeo Ichthyologica, 16, 1-32
The Lower Eocene (Ypresian) Chondrichthyes from Egem, Belgium. Part 1, Orders: Hexanchiformes, Squaliformes, Squatiniformes, Heterodontiformes and Orectolobiformes. Palaeo Ichthyologica, 15, 1–55