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The genus Heterodontus in the Early Paleogene of the North Sea Basin and the Landes, France. Palaeo Ichthyologica, 17, 1–60
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On the dentition of the extant species Heterodontus mexicanus and Heterodontus quoyi. Palaeo Ichthyologica, 16, 1-32
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The Lower Eocene (Ypresian) Chondrichthyes from Egem, Belgium. Part 1, Orders: Hexanchiformes, Squaliformes, Squatiniformes, Heterodontiformes and Orectolobiformes. Palaeo Ichthyologica, 15, 1–55