Nomura, M. & Hatanaka, O. & Nishimoto, H. & Karasawa, H. & Nanao Nojiriko Group (1991)
Megasqualus serriculus Jordan and Hannibal (Squalidae: Squaliformes: Elasmobranchii) from the Middle Miocene Nanao Calcareous Sandstone, Nanao City, Noto Peninsula, Central Japan. Bulletin of the Mizunami Fossil Museum, 18, 33–45