Maturity, spermatogenesis and seasonal reproductive cycle of male Rhinobatos annulatus (Muller & Henle, 1841) from Algoa Bay, South Africa, and a novel description for sperm release from the spermatocyst. African Zoology, 49(1), 128–136
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Spermatogenesis in the male white-spotted houndshark, Mustelus palumbes Smith, 1957. South African Journal of Science, 89(5), 244–246
Function of the liver and hepatic lipids of the lesser sand shark, Rhinobatos annulatus (Müller & Henle). Comparative Biochemistry and Physiology – Part B, Comparative Biochemistry, 86(4), 785–790
DOI: 10.1016/0305-0491(87)90225-2
Age and growth of the sand shark, Rhinobatos annulatus, in Algoa Bay, South Africa. Journal of Fish Biology, 25(2), 213–222
DOI: 10.1111/j.1095-8649.1984.tb04868.x