Cochran, J.E.M. & Charles, R. & Temple, A.J. & Kyne, P.M. & García‐Rodríguez, E. & Gonzalez‐Pestana, A. & Batlle‐Morera, A. & Mouton, T.L. & Armstrong, A.O. & Rohner, C.A. & Coker, D.J. & Hardenstine, R.S. & Kattan, A. & McIvor, A.J. & Peinemann, V.N. & O’Toole, K.A. & Palm, L. & Richardson, E.B. & Akhilesh, K.V. & Abedi, H.A. & Almealla, R.K. & Almojil, D. & Andrzejaczek, S. & Askin, A.N. & Banerjee, A.A. & Bargahi, H.R. & Barnes, A.J. & Barteneva-Vitry, S. & Behzadi, S. & Bein, A. & Bennett, R.H. & Bocchi, F. & Boldrocchi, G. & Braulik, G.T. & Braun, C.D. & Brighton, E. & Budd, F.K.P. & Bullock, R.W. & Canovas Perez, C. & Carlisle, A.B. & Carpenter, M. & Chapple, T.K. & Chaúca, I. & Cliff, G. & Crochelet, E. & Cullain, N. & Curnick, D.J. & Daly, R. & de Necker, L. & Diamant, S. & Donati, G.F.A. & Ebert, D.A. & Eid, E. & Elhassa, I.S. & Elston, C. & Everett, B.I. & Farrag, M.M.S. & Fassbender, N. & Fennessy, S.T. & Fernando, S.M.C. & Finucci, B. & Flam, A.L. & Gausman, P. & Gauthier, A.R.G. & Sreekanth, G.B. & Gupta, T. & Hafeez, M. & Hagy, B.N. & Haines, J.L.A. & Harris, J.L. & Harvey-Carroll, J. & Hempson, T.N. & Hilbourne, S.T. & Hsu, H.H. & Ibrahim, N.D. & Jacoby, D.M.P. & Jaquemet, S. & K K, I.B. & Karnad, D. & Kaunda-Arara, B. & Kizhakudan, S.J. & Kock, A.A. & Koester, A. & Kuboja, B.N. & Kuguru, B.L. & Lea, J.S.E. & Mahadalle, O. & Manjebrayakath, H. & Mason-Parker, C. & Mateos-Molina, D. & Menon, M. & Moore, A.B.M. & Mourier, J. & Murra, T.S. & Nakhawa, A.D. & Nazurally, N. & Nelso, L.E. & Nevill, J.E.G. & Olbers, J.M. & Ostrovski, R.L. & Peel, L.R. & Perisic, N. & Peterson, B. & Pierce, S.J. & Pittman, S.J. & Rahangdale, S. & Rambahiniarison, J. & Rastgoo, A.R. & Rezaie-Atagholipour, M. & Robinson, D.P. & Samoilys, M.A. & Sawers, T.J. & Scannell, B.J. & Schmidt, J.V. & Silva, I.M. & Silva, L. & Solonomenjanahary, J. & Spaet, J.L.Y. & Stevens, G.M.W. & Strike, E.M. & Thomas, S. & van Beuningen, D. & Venables, S.K. & Vossgaetter, L. & Weideli, O.C. & Williams, I.D. & Williams, C.T. & Willson, A.J. & Wilson, L. & Zareer, I.H. & Zerr, K.M. & Berumen, M.L. & Jabado, R.W. (2026)
Only One Percent of Important Shark and Ray Areas in the Western Indian Ocean Are Fully Protected From Fishing Pressure. Ecology and Evolution, 16, Article 1
DOI: 10.1002/ece3.72690

Sen, S. & Dash, G. & Kizhakudan, S.J. & Thomas, S. & Zacharia, P.U. & Ghosh, S. & Pradhan, R.K. & Das, M. & Dash, B. (2025)
Life history and stock status of the critically endangered Smoothback Guitarfish in the Northwestern Bay of Bengal. Journal of Coastal Conservation, 29(1), Article 19
DOI: 10.1007/s11852-025-01102-4

Wilson, L. & Roul, S.K. & Dash, S.S. & Remya, L. & Mahesh, V. & Akhilesh, K.V. & Rahangdale, S. & Purushottama, G.B. & Muktha, M. & Surya, S. & Najmudeen, T.M. & Nakhawa, A.D. & Chellappan, A. & Abdul Azeez, P. & Vinothkumar, R. & Vidhya, V. & Rajesh, K.M. & Ganga, U. & Thomas, S. & Kizhakudan, S.J. (2025)
Illustrated Guide To Protected Elasmobranchs In Indian Waters - With WPAA, IUCN, CITES & NDF Status. CMFRI Booklet Series No. 44/2025

Remya, L. & Thomas, S. & Muktha, M. & Kizhakudan, S.J. (2025)
Incidence of pregnant endangered devil rays (Elasmobranchii: Mobulidae) from the Gulf of Mannar and Palk Bay, southeast coast of India. Current Science, 129(4), 362–369
DOI: 10.18520/cs/v129/i4/362-369

Garzon, F. & Abel, G. & Burrett, I. & Exeter, O.M. & Hawkes, L.A. & Horton, T.W. & Lambrette, M. & Plaster, A. & Righton, D. & Rudd, H. & Rudd, J.L. & Stewart, B.D. & Thomas, S. & Thorburn, J. & Whittaker, P.S.M. & Witt, M.J. (2025)
Survival and recovery of three shark species in North-East Atlantic recreational fisheries. Ices Journal of Marine Science, 82(10), Article fsaf191
DOI: 10.1093/icesjms/fsaf191

Purushottama, G.B. & Thomas, S. & Kizhakudan, S.J. (2025)
Biology of the widenose guitarfish Glaucostegus obtusus (Muller and Henle,1841) (Rhinopristiformes: Glaucostegidae) from the eastern Arabian Sea. Indian Journal of Fisheries, 72(3), 45–57
DOI: 10.21077/ijf.2025.72.3.154488-05

Menon, M. & Kizhakudan, S.J. & Thomas, S. & Salim, S.S. & Zacharia, P.U. & Sen Dash, S. & Akhilesh, K.V. & Najmudeem, T.M. & Malayilethu, V. & Mahesh, V. & Gop, A.P. & Remya, L. & Manojkumar, P.P. & Rahangdale, S. & Purushottama, G.B. & Roul, S.K. & Pradhan, R.K. & Wilson, L. & Nair, R.J. & Patil, G. & Jamalabad, A. & Yousuf, K. & Radhakrishnan, M. & Sunil, K.T.S. (2024)
Shark consumption in India: An exploratory study on trends and awareness. Indian Journal of Fisheries, 71(4), 115–123
DOI: 10.21077/ijf.2024.71.4.143316-15

Dash, S.S. & Das, P.C. & Das, M. & Das, Ma. & Thomas, S. (2024)
Report on the Silky shark landings in Odisha. Marine Fisheries Information Service, Technical and Extension Series, 258, 34–35

Thomas, S. & Mahesh, V. & Kizhakudan, S.J. & Suresh Babu, P.P. & Ramudu, K.R. & Hussain, T. & Durgeka, N.M. (2024)
Protected guitarfish released by fishermen of Uttara Kannada District, Karnataka. Marine Fisheries Information Service, Technical and Extension Series, 260, 28–29

Muktha, M. & Cherian, J. & Renuka, V. & Maharshi, A. & Pattnaik, P. & Moshe, C. & Prabhakar, R.V.D. & Rao, T.N. & Thomas, S. & Kizhakudan, S.J. (2024)
Landings of Tiger shark along north Andhra Pradesh coast. Marine Fisheries Information Service, Technical and Extension Series, 261, 40–41

Sukumaran, S. & Sebastian, W. & Zacharia, P.U. & Kizhakudan, S.J. & Akhilesh, K.V. & Thomas, S. & Roul, S.K. & Muktha, M. & Sen, S.P. & Purushottama, G.B. & Rahangdale, S. & Bineesh, K.K. & Kishor, T.G. & Gopalakrishnan, A. (2023)
Population genetic research on the Spadenose shark, Scoliodon laticaudus (Chondrichthyes, Carcharhinidae), reveals the presence of two significantly differentiated clades along the Indian coast. Marine Biodiversity, 53(1), Article 16
DOI: 10.1007/s12526-022-01324-x

Akhilesh, K.V. & Kizhakudan, S.J. & Muktha, M. & Najmudeen, T.M. & Thomas, S. & Karnad, D. & Sutaria, D. & Fernandes, M. & Gupta, T. & Namboothri, N. & Patankar, V. & Dash, S.S. & Varghese, S.P. & Biju Kumar, A. & Barnes, A. & Bineesh, K.K. & John, S. & Gangal, M. & Manjebrayakath, H. & Malayilethu, V. & Tyabji, Z. & Malaika, V. & Sukumaran, S. & Purushottama, G.B. & Wilson, L. & Mahesh, V. & Nair, R.J. & Remya, L. & Rahangdale, S. & Manojkumar, P.P. & Sivakumar, K. & Vivekanandan, E. & Zacharia, P.U. & Gopalakrishnan, A. (2023)
Elasmobranch conservation, challenges and management strategy in India: recommendations from a national consultative meeting. Current Science, 124(3), 292–303
DOI: 10.18520/cs/v124/i3/292-303

Lalitha, R. & Thomas, S. & Kizhakudan, S.J. & Shanmugam, T. & Ayyakannu, G. & Muthayan, M. & Raja, R. & Vedhagiri, V. & Umayan, R. & Kalimuthu, S. & Murugesan, P. (2023)
Biological observations and new maximum reported sizes for two threatened rays: ornate eagle ray Aetomylaeus vespertilio and mangrove whipray Urogymnus granulatus from the Gulf of Mannar, India. Journal of the Marine Biological Association of the United Kingdom, 103, Article e90
DOI: 10.1017/s0025315423000796

Dash, S.S. & Kizhakudan, S.J. & Thomas, S. & Dash, G. & Das, M. & Ghosh, S. (2023)
Note on an albino spadenose shark, Scoliodon laticaudus from the north-east coast of India. Marine Fisheries Information Service, Technical and Extension Series, 255, 39–41

Sen, S. & Thomas, S. & Joe Kizhakudan, S. & Dash, G. & Pradhan, R.K. & Ghosh, S. & Das, M. & Dash, B. & Zacharia, P.U. (2022)
New observations of the endangered giant freshwater whipray, Urogymnus polylepis, provide further evidence for its distribution and breeding in the north-east coast of India. Journal of Fish Biology, 101(6), 1611–1616
DOI: 10.1111/jfb.15223

Purushottama, G.B. & Thomas, S. & Kizhakudan, S.J. & Zacharia, P.U. (2022)
Catch composition, reproductive biology and diet of the bowmouth guitarfish Rhina ancylostomus Bloch and Shneider, 1801 (Batoidea: Rhinidae) in the eastern Arabian Sea, India. Indian Journal of Fisheries, 69(3), 1–11
DOI: 10.21077/ijf.2022.69.3.117929-01

Thomas, S. & Muktha, M. & Sen, S. & Kizhakudan, S.J. & Akhilesh, K.V. & Purushottama, G.B. & Mahesh, V. & Rahangdale, S. & Zacharia, P.U. & Najmudeen, T.M. & Manojkumar, P.P. & Remya, L. & Wilson, L. & Roul, S.K. & Pradhan, R. & Seetha, P.K. & Yousuf, K. & Nataraja, G.D. (2021)
Status of the hammerhead shark (Carcharhiniformes: Sphyrnidae) fishery in Indian waters with observations on the biology of scalloped hammerhead Sphyrna lewini (Griffith & Smith, 1834). Aquatic Conservation, Marine and Freshwater Ecosystems, 31(11), 3072–3086
DOI: 10.1002/aqc.3686

Thomas, S. & Purushottama, G.B. & Nataraja, G.D. & Kizhakudan, S.J. (2020)
Fishery and biological characteristics of the spadenose shark Scoliodon laticaudus Muller & Henle, 1838 from the Eastern Arabian Sea. Regional Studies in Marine Science, 34, Article 101085
DOI: 10.1016/j.rsma.2020.101085

Kizhakudan, S.J. & Akhilesh, K.V. & Thomas, S. & Yousuf, K.S.S.M. & Sobhana, K.S. & Purushottama, G.B. & Menon, M. & Dash, S.S. & Manojkumar, P.P. & Nair, R.J. & Najmudeen, T.M. & Zacharia, P.U. (2018)
Field identification of batoids - a guide to Indian species. CMFRI Special Publication No.132, ISSN 0972-2351

Dudgeon, C.L. & Coulton, L. & Bone, R. & Ovenden, J.R. & Thomas, S. (2017)
Switch from sexual to parthenogenetic reproduction in a zebra shark. Scientific Reports, 7, Article 40537

Tulenko, F.J. & Massey, J.L. & Holmquist, E. & Kigundu, G. & Thomas, S. & Smith, S.M.E. & Mazan, S. & Davis, M.C. (2017)
Fin-fold development in paddlefish and catshark and implications for the evolution of the autopod. Proceedings of the Royal Society of London, Series B, 284(1855), Article 20162718
DOI: 10.1098/rspb.2016.2780

Kizhakudan, S.J. & Zacharia, P.U. & Thomas, S. & And Vivekanandan, E. & Muktha, M. (2015)
Guidance on National Plan of Action for Sharks in India. CMFRI Marine Fisheries Policy Series(2): 1–102

Veena, S. & Thomas, S. & Raje, S.G. & Durgekar, N.R. (2011)
Case of leucism in the spadenose shark, Scoliodon laticaudus (Müller and Henle, 1838) from Mangalore, Karnataka. Indian Journal of Fisheries, 58(1), 109–112

Thomas, S. & Raje, S.G. & Gowda, N.C. & Naik, A.A. & Kemparaju, S. (2007)
On the first record of Pig eye shark, Carcharhinus amboinensis (Muller & Henle, 1839) from Karnataka. Marine Fisheries Information Service, Technical and Extension Series, 194, 20–21

Thomas, S. & Raje, S.G. & Gowda, N.C. & Naik, A.R. & Kemparaju, S. (2007)
First record of Pig eye shark, Carcharhinus amboinensis (Muller & Henle, 1839) from Karnataka Marine Fisheries Information Service, Technical and Extension Series, 194, 20–21

Walsh, P.J. & Wood, C.M. & Perry, S.F. & Thomas, S. (1994)
Urea transport of hepatocytes and red blood cells of selected elasmobranch and teleost fishes. Journal of Experimental Biology, 193, 321–335