Sukumaran, S. & Sebastian, W. & Zacharia, P.U. & Kizhakudan, S.J. & Akhilesh, K.V. & Thomas, S. & Roul, S.K. & Muktha, M. & Sen, S.P. & Purushottama, G.B. & Rahangdale, S. & Bineesh, K.K. & Kishor, T.G. & Gopalakrishnan, A. (2023)
Population genetic research on the Spadenose shark, Scoliodon laticaudus (Chondrichthyes, Carcharhinidae), reveals the presence of two significantly differentiated clades along the Indian coast. Marine Biodiversity, 53(1), Article 16
DOI: 10.1007/s12526-022-01324-x

Akhilesh, K.V. & Kizhakudan, S.J. & Muktha, M. & Najmudeen, T.M. & Thomas, S. & Karnad, D. & Sutaria, D. & Fernandes, M. & Gupta, T. & Namboothri, N. & Patankar, V. & Dash, S.S. & Varghese, S.P. & Biju Kumar, A. & Barnes, A. & Bineesh, K.K. & John, S. & Gangal, M. & Manjebrayakath, H. & Malayilethu, V. & Tyabji, Z. & Malaika, V. & Sukumaran, S. & Purushottama, G.B. & Wilson, L. & Mahesh, V. & Nair, R.J. & Remya, L. & Rahangdale, S. & Manojkumar, P.P. & Sivakumar, K. & Vivekanandan, E. & Zacharia, P.U. & Gopalakrishnan, A. (2023)
Elasmobranch conservation, challenges and management strategy in India: recommendations from a national consultative meeting. Current Science, 124(3), 292–303
DOI: 10.18520/cs/v124/i3/292-303

Lalitha, R. & Thomas, S. & Kizhakudan, S.J. & Shanmugam, T. & Ayyakannu, G. & Muthayan, M. & Raja, R. & Vedhagiri, V. & Umayan, R. & Kalimuthu, S. & Murugesan, P. (2023)
Biological observations and new maximum reported sizes for two threatened rays: ornate eagle ray Aetomylaeus vespertilio and mangrove whipray Urogymnus granulatus from the Gulf of Mannar, India. Journal of the Marine Biological Association of the United Kingdom, 103, Article e90
DOI: 10.1017/s0025315423000796

Sen, S. & Thomas, S. & Joe Kizhakudan, S. & Dash, G. & Pradhan, R.K. & Ghosh, S. & Das, M. & Dash, B. & Zacharia, P.U. (2022)
New observations of the endangered giant freshwater whipray, Urogymnus polylepis, provide further evidence for its distribution and breeding in the north-east coast of India. Journal of Fish Biology, 101(6), 1611–1616
DOI: 10.1111/jfb.15223

Purushottama, G.B. & Thomas, S. & Kizhakudan, S.J. & Zacharia, P.U. (2022)
Catch composition, reproductive biology and diet of the bowmouth guitarfish Rhina ancylostomus Bloch and Shneider, 1801 (Batoidea: Rhinidae) in the eastern Arabian Sea, India. Indian Journal of Fisheries, 69(3), 1–11
DOI: 10.21077/ijf.2022.69.3.117929-01

Thomas, S. & Muktha, M. & Sen, S. & Kizhakudan, S.J. & Akhilesh, K.V. & Purushottama, G.B. & Mahesh, V. & Rahangdale, S. & Zacharia, P.U. & Najmudeen, T.M. & Manojkumar, P.P. & Remya, L. & Wilson, L. & Roul, S.K. & Pradhan, R. & Seetha, P.K. & Yousuf, K. & Nataraja, G.D. (2021)
Status of the hammerhead shark (Carcharhiniformes: Sphyrnidae) fishery in Indian waters with observations on the biology of scalloped hammerhead Sphyrna lewini (Griffith & Smith, 1834). Aquatic Conservation, Marine and Freshwater Ecosystems, 31(11), 3072–3086
DOI: 10.1002/aqc.3686

Thomas, S. & Purushottama, G.B. & Nataraja, G.D. & Kizhakudan, S.J. (2020)
Fishery and biological characteristics of the spadenose shark Scoliodon laticaudus Muller & Henle, 1838 from the Eastern Arabian Sea. Regional Studies in Marine Science, 34, Article 101085
DOI: 10.1016/j.rsma.2020.101085

Kizhakudan, S.J. & Akhilesh, K.V. & Thomas, S. & Yousuf, K.S.S.M. & Sobhana, K.S. & Purushottama, G.B. & Menon, M. & Dash, S.S. & Manojkumar, P.P. & Nair, R.J. & Najmudeen, T.M. & Zacharia, P.U. (2018)
Field identification of batoids - a guide to Indian species. CMFRI Special Publication No.132, ISSN 0972-2351

Dudgeon, C.L. & Coulton, L. & Bone, R. & Ovenden, J.R. & Thomas, S. (2017)
Switch from sexual to parthenogenetic reproduction in a zebra shark. Scientific Reports, 7, Article 40537

Tulenko, F.J. & Massey, J.L. & Holmquist, E. & Kigundu, G. & Thomas, S. & Smith, S.M.E. & Mazan, S. & Davis, M.C. (2017)
Fin-fold development in paddlefish and catshark and implications for the evolution of the autopod. Proceedings of the Royal Society of London, Series B, 284(1855), Article 20162718
DOI: 10.1098/rspb.2016.2780

Kizhakudan, S.J. & Zacharia, P.U. & Thomas, S. & And Vivekanandan, E. & Muktha, M. (2015)
Guidance on National Plan of Action for Sharks in India. CMFRI Marine Fisheries Policy Series(2): 1–102

Veena, S. & Thomas, S. & Raje, S.G. & Durgekar, N.R. (2011)
Case of leucism in the spadenose shark, Scoliodon laticaudus (Müller and Henle, 1838) from Mangalore, Karnataka. Indian Journal of Fisheries, 58(1), 109–112

Thomas, S. & Raje, S.G. & Gowda, N.C. & Naik, A.A. & Kemparaju, S. (2007)
On the first record of Pig eye shark, Carcharhinus amboinensis (Muller & Henle, 1839) from Karnataka. Marine Fisheries Information Service, Technical and Extension Series, 194, 20–21

Walsh, P.J. & Wood, C.M. & Perry, S.F. & Thomas, S. (1994)
Urea transport of hepatocytes and red blood cells of selected elasmobranch and teleost fishes. Journal of Experimental Biology, 193, 321–335