Azofeifa-Solano, J.C. & Clarke, T.M. & Espinoza, M. & Wehrtmann, I.S. (2021)
Reproductive cycles of the southern banded guitarfish Zapteryx xyster and the Velez ray Raja velezi caught as bycatch in a bottom-trawl fishery. Latin American Journal of Aquatic Research, 49(1), 48–59
DOI: 10.3856/vol49-issue1-fulltext-2558

Clarke, T.M. & Espinoza, M. & Chaves, R.R. & Wehrtmann, I.S. (2017)
Assessing the vulnerability of demersal elasmobranchs to a data-poor shrimp trawl fishery in Costa Rica, Eastern Tropical Pacific. Biological Conservation, 217, 321–328
DOI: 10.1016/j.biocon.2017.11.015

Sandoval-Herrera, N.I. & Vargas-Soto, J.S. & Espinoza, M. & Clarke, T.M. & Fisk, A.T. & Wehrtmann, I.S. (2016)
Mercury levels in muscle tissue of four common elasmobranch species from the Pacific coast of Costa Rica, Central America. Regional Studies in Marine Science, 3, 254–261
DOI: 10.1016/j.rsma.2015.11.011

Espinoza, M. & Munroe, S.E.M. & Clarke, T.M. & Fisk, A.T. & Wehrtmann, I.S. (2015)
Feeding ecology of common demersal elasmobranch species in the Pacific coast of Costa Rica inferred from stable isotope and stomach content analyses. Journal of Experimental Marine Biology and Ecology, 470, 12–25
DOI: 10.1016/j.jembe.2015.04.021

Clarke, T.M. & Espinoza, M. & Ahrens, R. & Wehrtmann, I.S. (2015)
Elasmobranch bycatch associated with the shrimp trawl fishery off the Pacific coast of Costa Rica, Central America. Fishery Bulletin, 114(1), 1–17
DOI: 10.7755/FB.114.1

Clarke, T.M. & Espinoza, M. & Wehrtmann, I.S. (2014)
Reproductive ecology of demersal elasmobranchs from a data-deficient fishery, Pacific of Costa Rica, Central America. Fisheries Research, 157, 96–105
DOI: 10.1016/j.fishres.2014.04.003

Espinoza, M. & Clarke, T.M. & Villalobos-Rojas, F. & Wehrtmann, I.S. (2013)
Diet composition and diel feeding behaviour of the banded guitarfish Zapteryx xyster along the Pacific coast of Costa Rica, Central America. Journal of Fish Biology, 82(1), 286–305
DOI: 10.1111/j.1095-8649.2012.03488.x

Espinoza, M. & Clarke, T. & Villalobos-Rojas, F. & Wehrtmann, I.S. (2012)
Dietary overlap and resource partitioning of four elasmobranch species off the Pacific of Costa Rica: implications for management [Abstract]. World Congress of Herpetology, Vancouver, Canada, August 8 – 14, 2012, Book of Abstracts, 199–200

Espinoza, M. & Clarke, T.M. & Villalobos-Rojas, F. & Wehrtmann, I.S. (2012)
Ontogenetic dietary shifts and feeding ecology of the rasptail skate Raja velezi and the brown smoothhound shark Mustelus henlei along the Pacific coast of Costa Rica, Central America. Journal of Fish Biology, 81(5), 1517–1595
DOI: 10.1111/j.1095-8649.2012.03410.x

Clarke, T.M. & Espinoza, M. & Villalobos-Rojas, F. & Wehrtmann, I.S. (2012)
Reproductive ecology of four elasmobranch species in the Pacific coast of Costa Rica, Central America [Abstract]. World Congress of Herpetology, Vancouver, Canada, August 8 – 14, 2012, Book of Abstracts, 136

Clarke, T.M. & Espinoza, M. & Villalobos-Rojas, F. & Wehrtmann, I.S. (2011)
Summary of Demersal Elasmobranch Studies in the Continental Platform of the Pacific of Costa Rica with Management and Conservation Strategies. Technical Report. San José, Costa Rica