Bourdon, J. (1999)
A fossil manta from the Early Pliocène (Zanclean) of North America. Tertiary Research, 19(3–4), 79–84
Adnet, S. & Cappetta, H. & Guinot, G. & Notarbartolo Di Sciara, G. (2012)
Evolutionary history of the devilrays (Chondrichthyes: Myliobatiformes) from fossil and morphological inference. Zoological Journal of the Linnean Society, 166(1), 132–159
DOI: 10.1111/j.1096-3642.2012.00844.x