Smith, A. (1838)
Pisces. Illustrations of the zoology of South Africa. Smith, Elder, London, vol. 4, 77 pp., not numbered, pls 1–31

Müller, J. & Henle, F.G.J. (1841)
Systematische Beschreibung der Plagiostomen. Berlin, Veit, pp. 1–200

Gray, J.E. (1851)
List of the specimens of fish in the collection of the British Museum. Part I. Chondropterygii. British Museum (Natural History), London, 160 pp

Pappe, L. (1853)
Synopsis of the edible fishes at the Cape of Good Hope. Cape Town.

Dumeril, A.H.A. (1865)
Histoire naturelle des poissons ou ichthyologie générale. Tome Premier. Elasmobranchés. Plagiostomes et Holocéphales ou Chimères. Librairie Encyclopédique de Roret, Paris. 720 pp

Tiesing, B. (1895)
Ein Beitrag zur Kenntnis der Augen-, Kiefer- und Kiemenmuskulatur der Haie und Rochen. Jenaische Zeitschrift für Naturwissenschaft, 30 (n.F. Bd. 23), 75–126

Jordan, D.S. & Snyder, J.O. (1901)
A preliminary check-list of the fishes of Japan. Annotationes Zoologicae Japonenses, 3, 31–160

Garman, S. (1913)
The Plagiostomia (Sharks, Skates and Rays). Memoirs of the Museum of Comparative Zoology at Harvard College, 36, 1–528

Pellegrin, J. (1914)
Sur une collection de poissons de Madagascar. Bulletin de la Société zoologique de France, 39, 221–234

Barnard, K.H. (1925)
A monograph of the marine fishes of South Africa. Part I (Amphioxus, Cyclostomata, Elasmobranchii, and Teleostei -Isospondyli to Heterosomata). Annals of the South African Museum, 21(1), 1–418

Norman, J.R. (1926)
A synopsis of the rays of the family Rhinobatidae, with a revision of the genus Rhinobatus. Proceedings of the Zoological Society of London, 96(4), 941–982
DOI: 10.1111/j.1096-3642.1926.tb02228.x

Fowler, H.W. (1941)
Contributions to the biology of the Philippine archipelago and adjacent regions. The fishes of the groups Elasmocephalii, Holocephali, Isospondyli, and Ostariophysi obtained by the United States Fisheries Steamer "Albatross" in 1907 to 1910, chiefly in the Philippine islands and adjacent seas. Bulletin of the United States National Museum, 100, i–x, 1–879

Smith, J.L.B. (1953)
The sea fishes of Southern Africa. Revised enlarged edition. Johannesburg: 1–564, Pls. 1–107.

Smith, J.L.B. (1961)
The sea fishes of Southern Africa. Johannesburg Central News Agency Ltd., 573 p. 4th ed.

Fourmanoir, P. (1963)
Raies et requins-scie de la côte ouest de Madagascar (Ordre des Batoidei). Cahiers Orstom, Série Océanographie, 1963, (6), 33–58

Kiener, A. (1965)
Contribution à l’étude écologique et biologique des eaux saumâtres malgaches. Les poissons euryhalins et leur rôle dans le développement des pêches. Vie et Milieu, Serie C, 16(2), 1013–1150

Wallace, J.H. (1967)
The batoid fishes of the east coast of southern Africa. Part I: Sawfishes and guitarfishes. Investigational Report Oceanographic Research Institute, 15, 1–32

Rossouw, G.J. (1984)
Age and growth of the sand shark, Rhinobatos annulatus, in Algoa Bay, South Africa. Journal of Fish Biology, 25(2), 213–222
DOI: 10.1111/j.1095-8649.1984.tb04868.x

Compagno, L.J.V. (1986)
Families Pristidae, Torpedinidae, Narkidae, Rhinobatidae, Myliobatidae, Mobulidae, Dasyatidae, Callorhinchidae, Chimaeridae, Rhinochimaeridae In Smith's Sea Fishes, by M.M. Smith & P.C. Heemstra, eds. Macmillian, Johannesburg: 110–114, 128–142, 144–148

Smith, M.M. & Heemstra, P.C. (1986)
Smiths' Sea fishes Braamfontein, Johannesburg :Macmillan South Africa,1986.

Rossouw, G.J. (1987)
Function of the liver and hepatic lipids of the lesser sand shark, Rhinobatos annulatus (Müller & Henle). Comparative Biochemistry and Physiology – Part B, Comparative Biochemistry, 86(4), 785–790
DOI: 10.1016/0305-0491(87)90225-2

Du Preez, H.H. & Mclachlan, A. & Marais, J.F.K. (1988)
Oxygen consumption of two nearshore marine elasmobranchs, Rhinobatos annulatus (Müller & Henle, 1841) and Myliobatus aquila (Linnaeus, 1758). Comparative Biochemistry and Physiology – Part A, Physiology, 89(2), 283–294
DOI: 10.1016/0300-9629(88)91094-8

Harris, S.A. & Bennett, B.A. & Branch, G.M. (1988)
An assessment of the role of the sand shark Rhinobatos annulatus as a predator in Langebaan Lagoon. South African Journal of Marine Science, 7(1), 153–159
DOI: 10.2989/025776188784379125

Compagno, L.J.V. & Ebert, D.A. & Smale, M.J. (1989)
Guide to the sharks and rays of Southern Africa. New Holland Ltd., London 1989

Du Preez, H.H. & Mclachlan, A. & Marais, J.F.K. & Cockcroft, A.C. (1990)
Bioenergetics of fishes in a high-energy surf-zone. Marine Biology, 106(1), 40513
DOI: 10.1007/BF02114669

Compagno, L.J.V. & Ebert, D.A. & Cowley, P.D. (1991)
Distribution of offshore demersal cartilaginous fishes (Class Chondrichthyes) of the west coast of southern Africa, with notes on their systematics. South African Journal of Marine Science, 11(1), 43–139, figs 1–24
DOI: 10.2989/025776191784287664

Deynat, P.P. & Séret, B. (1996)
Le revêtement cutané des raies (Chondrichthyes, Elasmobranchii, Batoidea). I- Morphologie et arrangement des denticules cutanés. Annales des Sciences Naturelles – Zoologie et Biologie Animale, 17(2), 65–83

Van As, J.G. & Basson, L. (1996)
An endosymbiotic trichodinid, Trichodina rhinobatae sp n (Ciliophora: Peritrichia) found in the lesser guitarfish, Rhinobatos annulatus Smith, 1841 (Rajiformes: Rhinobatidae) from the South African coast. Acta Protozoologica, 35(1), 61–67

Clark, B.M. (1997)
Variation in surf-zone fish community structure across a wave-exposure gradient. Estuarine, Coastal and Shelf Science, 44(6), 659–674
DOI: 10.1006/ecss.1996.0151

Mould, B. (1997)
Classification of the recent Elasmobranchii. Copyright Brian Mould 1997

Deynat, P.P. (1998)
Le revêtement cutané des raies (Chondrichthyes, Elasmobranchii, Batoidea). II. Morphologie et arrangement des tubercules cutanés. Annales des Sciences Naturelles – Zoologie et Biologie Animale, 19(3–4), 155–172
DOI: 10.1016/S0003-4339(99)80004-5

Bianchi, G. & Carpenter, K.E. & Roux, J.-P. & Molloy, F.J. & Boyer, D. & Boyer, H.J. (1999)
FAO species identification field guide for fishery purposes. The living marine resources of Namibia. FAO, Rome. 250 p.

Compagno, L.J.V. (1999)
An overview of chondrichthyan systematics and biodiversity in southern Africa. Transactions of the Royal Society of South Africa, 54, 75–120
DOI: 10.1080/00359199909520406

Compagno, L.J.V. (2005)
Checklist of living Chondrichthyes. In W.C. Hamlett (Ed.), Reproductive biology and phylogeny of chondrichthyes: sharks, rays and chimaeras, Vol. 3 (pp. 503–548). Science Publishers, Endfield, USA

Burgess, G.H. & Holtzhausen, H.A. & Smale, M.J. (2006)
Rhinobatos annulatus IUCN 2012, IUCN Red List of Threatened Species, Version 2012.2, https://www.iucnredlist.org

Lamberth, S.J. (2006)
White shark and other chondrichthyan interactions with the beach-seine (treknet) fishery in False Bay, South Africa. African Journal of Marine Science, 28(3–4), 723–727
DOI: 10.2989/18142320609504222

Pradervand, P. & Hiseman, R. (2006)
An analysis of the recreational shore fishery in the Goukamma Marine Protected Area. African Zoology, 41(2), 275–289
DOI: 10.3377/1562-7020(2006)41[275:aaotrs]2.0.co;2

Pradervand, F. & Mann, B.Q. & Bellis, M.F. (2007)
Long-term trends in the competitive shore fishery along the KwaZulu-Natal coast, South Africa. African Zoology, 42(2), 216–236
DOI: 10.3377/1562-7020(2007)42[216:LTITCS]2.0.CO;2

da Silva, C. & Bürgener, M. (2007)
South Africa's demersal shark meat harvest. Traffic Bulletin, 21(2), 55–66

Béarez, P. & Kerneur, S. & Gabsi, Z. (2008)
New record of a surgeonfish from Oman with notes on some uncommon rays. Cybium, 32(4), 355–358

Dicken, M.L. (2010)
The ichthyofauna in the Port of Ngqura, South Africa. African Journal of Marine Science, 32(3), 491–499
DOI: 10.2989/1814232x.2010.538139

Vaughan, D.B. & Chisholm, L.A. (2010)
A new species of Neoheterocotyle Hargis, 1955 (Monogenea: Monocotylidae) from the gills of Rhinobatos annulatus Müller & Henle (Rhinobatidae) off the southern tip of Africa. Systematic Parasitology, 77(3), 205–213
DOI: 10.1007/s11230-010-9268-5

Attwood, C.G. & Petersen, S.L. & Kerwath, S.E. (2011)
Bycatch in South Africa's inshore trawl fishery as determined from observer records. ICES Journal of Marine Science, 68(10), 2163–2174
DOI: 10.1093/icesjms/fsr162

Vaughan, D.B. & Chisholm, L.A. (2011)
Amendment of Pseudoleptobothrium Young, 1967 (Monogenea, Microbothriidae) with the description of Pseudoleptobothrium christisoni sp. nov. from the dermal denticles of Rhinobatos annulatus (Rhinobatidae) off the southern tip of Africa. Acta Parasitologica, 56(3), 280–289
DOI: 10.2478/s11686-011-0057-3

Naylor, G.J.P. & Caira, J.N. & Jensen, K. & Rosana, K.A.M. & White, W.T. & Last, P.R. (2012)
A DNA sequence based approach to the identification of shark and ray species and its implications for global elasmobranch diversity and parasitology. Bulletin of the American Museum of Natural History, 367, 1–262

Caira, J.N. & Marques, F.P.L. & Jensen, K. & Kuchta, R. & Ivanov, V. (2013)
Phylogenetic analysis and reconfiguration of genera in the cestode order Diphyllidea. International Journal for Parasitology, 43(8), 621–639
DOI: 10.1016/j.ijpara.2013.03.001

Caira, J.N. & Pickering, M. & Schulman, A.D. & Hanessian, N.J. (2013)
Two New Species of Echinobothrium (Cestoda: Diphyllidea) from Batoids off South Africa. Comparative Parasitology, 80(1), 22–32

Akhilesh, K.V. & Bineesh, K.K. & Gopalakrishnan, A. & Jena, J.K. & Basheer, V.S. & Pillai, N.G.K. (2014)
Checklist of Chondrichthyans in Indian waters. Journal of the Marine Biological Association of India, 56(1), 109–120
DOI: 10.6024/jmbai.2014.56.1.01750s-17

Rossouw, G.J. (2014)
Maturity, spermatogenesis and seasonal reproductive cycle of male Rhinobatos annulatus (Muller & Henle, 1841) from Algoa Bay, South Africa, and a novel description for sperm release from the spermatocyst. African Zoology, 49(1), 128–136
DOI: 10.3377/004.049.0116

Ebert, D.A. & Van Hees, K.E. (2015)
Beyond Jaws: rediscovering the ‘lost sharks’ of southern Africa. African Journal of Marine Science, 37(2), 141–156
DOI: 10.2989/1814232X.2015.1048730

Ebert, D.A. & Van Hees, K.E. (2015)
Corrigendum: Beyond Jaws: rediscovering the ‘lost sharks’ of southern Africa. African Journal of Marine Science, 37(3), 435
DOI: 10.2989/1814232X.2015.1097052

Jones, C.M. & Driggers, W.B. & Castro, J.I. & de Carvalho, M.R. (2015)
On the attribution of authorship for several elasmobranch species in Müller and Henle’s Systematische Beschreibung der Plagiostomen (Chondrichthyes, Elasmobranchii). Zootaxa, 4052(5), 569–572
DOI: 10.11646/zootaxa.4052.5.4

de Vosa, L. & Watson, R.G.A. & Götz, A. & Attwood, C.G. (2015)
Baited remote underwater video system (BRUVs) survey of chondrichthyan diversity in False Bay, South Africa. African Journal of Marine Science, 37(2), 209–218
DOI: 10.2989/1814232X.2015.1036119

Last, P.R. & Séret, B. & Naylor, G.J.P. (2016)
A new species of guitarfish, Rhinobatos borneensis sp. nov. with a redefinition of the family-level classification in the order Rhinopristiformes (Chondrichthyes: Batoidea). Zootaxa, 4117(4), 451–475
DOI: 10.11646/zootaxa.4117.4.1

Morris, T. & Avenant-Oldewage, A. & Lamberth, S. & Reed, C. (2016)
Shark parasites as bio-indicators of metals in two South African embayments. Marine Pollution Bulletin, 104(1–2), 221–228
DOI: 10.1016/j.marpolbul.2016.01.027

Séret, B. & Last, P.R. & Naylor, G.J.P. (2016)
Guitarfishes, Family Rhinobatidae. In Last, P.R., White, W.T., Carvalho, M.R. de, Séret, B., Stehmann, M.F.W & Naylor, G.J.P(Eds.) Rays of the World. CSIRO Publishing, Melbourne: 78–109

Weigmann, S. (2016)
Annotated checklist of the living sharks, batoids and chimaeras (Chondrichthyes) of the world, with a focus on biogeographical diversity. Journal of Fish Biology, 88(3), 837–1037
DOI: 10.1111/jfb.12874

Moore, A.B.M. (2017)
Are guitarfishes the next sawfishes? Extinction risk and an urgent call for conservation action. Endangered Species Research, 34, 75–88
DOI: 10.3354/esr00830

Buckley, K.A. & Crook, D.A. & Pillans, R.D. & Smith, L. & Kyne, P.M. (2018)
Sustainability of threatened species displayed in public aquaria, with a case study of Australian sharks and rays. Reviews in Fish Biology and Fisheries, 28(1), 137–151
DOI: 10.1007/s11160-017-9501-2

Almeida, A.J. & Biscoito, M. (2019)
Keys to the fishes of the Eastern Atlantic Ocean, Mediterranean Sea and Black Sea. I – Myxini; Petromyzontida; Chondrichthyes Boletim do Museu de História Natural do Funchal, Suppl. 15, 1–195

Osgood, G.J. & McCord, M.E. & Baum, J.K. (2019)
Using baited remote underwater videos (BRUVs) to characterize chondrichthyan communities in a global biodiversity hotspot. PLoS ONE, 14(12), Article e0225859
DOI: 10.1371/journal.pone.0225859

Schaeffner, B.C. & Smit, N.J. (2019)
Parasites of cartilaginous fishes (Chondrichthyes) in South Africa - a neglected field of marine science. Folia Parasitologica, 66, Article 002
DOI: 10.14411/fp.2019.002

d'Alberto, B.M. & Carlson, J.K. & Pardo, S.A. & Simpfendorfer, C.A. (2019)
Population productivity of shovelnose rays: Inferring the potential for re-covery. PLoS ONE, 14(11), Article e0225183
DOI: 10.1371/journal.pone.0225183

Guinot, G. & Cavin, L. (2020)
Distinct Responses of Elasmobranchs and Ray-Finned Fishes to Long-Term Global Change. Frontiers in Ecololgy and Evolution, 7, Article 513
DOI: 10.3389/fevo.2019.00513

Osgood, G.J. & McCord, M.E. & Baum, J.K. (2020)
Chondrichthyans as an umbrella species-complex for conserving South African biodiversity. African Journal of Marine Science, 42(1), 81–93
DOI: 10.2989/1814232x.2020.1729859

Ebert, D.A. & Wintner, S.P. & Kyne, P.M. (2021)
An annotated checklist of the chondrichthyans of South Africa. Zootaxa, 4947(1), 1–127
DOI: 10.11646/zootaxa.4947.1.1

Augustine, S. & Lika, K. & Kooijman, S. (2022)
The comparative energetics of the chondrichthyans reveals universal links between respiration, reproduction and lifespan. Journal of Sea Research, 185, Article 102228
DOI: 10.1016/j.seares.2022.102228

Heemstra, P.C. & Heemstra, E. & Ebert, D.A. & Holleman, W. & Randall, J.E. (2022)
Coastal Fishes of the Western Indian Ocean. South African Institute for Aquatic Biodiversity, Private Bag 1015, Makhanda, 6140, South Africa

van Staden, M. & Ebert, D.A. & da Silva, C. & Bester-van der Merwe, A.E. (2022)
Comparative analyses of the complete mitochondrial genomes of two southern African endemic guitarfish, Acroteriobatus annulatus and A. blochii. International Journal of Biological Macromolecules, 223, 1094–1106
DOI: 10.1016/j.ijbiomac.2022.10.285

Barrowclift, E. & Gravel, S.M. & Pardo, S.A. & Bigman, J.S. & Berggren, P. & Dulvy, N.K. (2023)
Tropical rays are intrinsically more sensitive to overfishing than the temperate skates. Biological Conservation, 281, Article 110003
DOI: 10.1016/j.biocon.2023.110003

Faure-Beaulieu, N. & Lombard, A.T. & Olbers, J. & Goodall, V. & da Silva, C. & Daly, R. & Jordaan, G. & Kerwath, S.E. & Kock, A. & Mann, B.Q. & Murray, T.S. & Albano, P. & Cliff, G. & dos Santos, N.A. & Gennari, E. & Hammerschlag, N. & van der Merwe, A.E.B. & Watson, R. & Andreotti, S. & Bernard, A.T.F. & Cowley, P.D. & De Vos, L. & Drobniewska, N. & Elston, C. & Fallows, C. & Rogers, T.D. & Smith, G. & van Staden, M. & de Villiers, P. & Paulet, T.G. & Harris, J. (2023)
A systematic conservation plan identifying critical areas for improved chondrichthyan protection in South Africa. Biological Conservation, 284, Article 110163
DOI: 10.1016/j.biocon.2023.110163

Murray, T.S. & Elston, C. & Bennett, R.H. & Childs, A.R. & Cowley, P.D. (2023)
Movement patterns and underestimation of the maximum age of a Vulnerable endemic guitarfish species inferred from mark-recapture studies. African Journal of Marine Science, 45(2), 149–154
DOI: 10.2989/1814232x.2023.2224829

Reinecke, T. & Mollen, F.H. & Seitz, J.C. & Motomura, H. & Hovestadt, D. & Hoedemakers, K. (2023)
Iconography of jaws and representative teeth of extant rhinopristiform and dasyatoid batoids (Chondrichthyes, Elasmobranchii) for comparison with fossil batoid material. Palaeontos, 34, 3–158

Sherman, C.S. & Simpfendorfer, C.A. & Haque, A.B. & Digel, E.D. & Zubick, P. & Eged, J. & Matsushiba, J.H. & Sant, G. & Dulvy, N.K. (2023)
Guitarfishes are plucked: Undermanaged in global fisheries despite declining populations and high volume of unreported international trade. Marine Policy, 155, Article 105753
DOI: 10.1016/j.marpol.2023.105753

Elston, C. & Murray, T.S. (2024)
Multi-method approach identifies a South African estuary as an important elasmobranch habitat and potential nursery ground. African Journal of Marine Science, in press
DOI: 10.2989/1814232x.2024.2326814

Giareta, E.P. & Leite, R.D. & Davis, R.A.H. & Chaves, A.P. & Charvet, P. & Wosnick, N. (2024)
Unveiling the batoid plight: insights from global stranding data and future directions. Reviews in Fish Biology and Fisheries, in press
DOI: 10.1007/s11160-024-09837-3

Pollom, R.A. & Cheok, J. & Pacoureau, N. & Gledhill, K.S. & Kyne, P.M. & Ebert, D.A. & Jabado, R.W. & Herman, K.B. & Bennett, R.H. & da Silva, C. & Fernando, S. & Kuguru, B. & Leslie, R.W. & McCord, M.E. & Samoilys, M. & Winker, H. & Fennessy, S.T. & Pollock, C.M. & Rigby, C.L. & Dulvy, N.K. (2024)
Overfishing and climate change elevate extinction risk of endemic sharks and rays in the southwest Indian Ocean hotspot. Plos One, 19(9), Article e0306813
DOI: 10.1371/journal.pone.0306813

Pytka, J.M. & Kyne, P.M. & Carlson, J.K. & Wosnick, N. & Jabado, R.W. (2024)
A tangled web: global review of fishing interactions with rhino rays. Reviews in Fish Biology and Fisheries, 34(1), 131–160
DOI: 10.1007/s11160-023-09821-3

Vaughan, D.B. & Hansen, H. & Chisholm, L.A. (2024)
Proposal of Troglocephalinae n. subfam. (Monogenea: Monocotylidae) to accommodate existing and two new monocotylids from the gills of rhinopristiform shovelnose rays. Systematic Parasitology, 101, Article 51
DOI: 10.1007/s11230-024-10174-z