Additions to the fish fauna of the English Palaeogene. 1. Two new species of Alopias (Thresher Shark) from the English Eocene. Tertiary Research, 2(1): 23–28, 6 fig.
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London Clay Fossils of Kent and Essex. Rochester, Kent, Medway Fossil and Mineral Society, 228 p, ISBN: 978–0–9538243–1–1
A study of the sharks and rays from the Lillebælt Clay (Early–Middle Eocene) of Denmark, and their palaeoecology. Bulletin of the Geological Society of Denmark, 62: 39–88
Climate cooling and clade competition likely drove the decline of lamniform sharks. Proceedings of the National Academy of Sciences of the United States of America, 116(41), 20584–20590