de Carvalho, M.R. & Maisey, J.G. & Grande, L. (2004)
Freshwater stingrays of the Green River Formation of Wyoming (Early Eocene), with the description of a new genus and species and an analysis of its phylogenetic relationships (Chondrichthyes: Myliobatiformes). Bulletin of the American Museum of Natural History, 284, 1–136, 53 fig., 7 tabl.

Villalobos-Segura, E. & Underwood, C.J. (2020)
Radiation and divergence times of Batoidea. Journal of Vertebrate Paleontology, 40(3), Article e1777147
DOI: 10.1080/02724634.2020.1777147

López-Romero, F.A. & Villalobos-Segura, E. & Tuertscher, J. & Berio, F. & Stumpf, S. & Dearden, R.P. & Kriwet, J. & Maldonado, E. (2025)
Evolution of the batoidea pectoral fin skeleton: convergence, modularity, and integration driving disparity trends. Evolutionary Ecology, 39(1), 111–134
DOI: 10.1007/s10682-025-10330-x