Séret, B. (1986)
Deep Water Skates from Madagascar (Chondrichthyes, Rajoidei). In Uyeno, T. & Arai, R. & Taniuchi, T. & Matsuura, K. (Eds) Proceedings of the second international conference on Indo-Pacific fishes. Ichthyological Society of Japan, Tokyo: 256–260

Ishihara, H. & Treloar, M. & Bor, P.H.F. & Senou, H. & Jeong, C.-H. (2012)
The Comparative Morphology of Skate Egg Capsules (Chondrichthyes: Elasmobranchii: Rajiformes). Bulletin of the Kanagawa Prefectural Museum, Natural Science, 41, 9–25