Jaekel, O. (1899)
Ueber die Organisation der Petalodonten. Zeitschrift der Deutschen Geologischen Gesellschaft, 51, 258–298

Lebedev, O.A. & Esin, D. (2007)
Palaeopathological evidence in some permocarboniferous fishes. Ichthyolith Issues, Special Publication, 10, 59–60

Ginter, M. & Hampe, O. & Duffin, C.J. (2010)
Handbook of Paleoichthyology, Vol. 3D: Chondrichthyes Paleozoic Elasmobranchii: Teeth Verlag Dr. Friedrich Pfeil, pp. 168, 154 fig., 4 Tab.