Jordan, D.S. & Evermann, B.W. (1898)
The fishes of North and Middle America: a descriptive catalogue of the species of fish-Iike vertebrates found in the waters of North America, north of the Isthmus of Panama. Part III. Bulletin of the United States National Museum, 47(3), 2183a–3136
Gilbert, C.H. & Starks, E.C. (1904)
The fishes of Panama Bay. Memoirs of the California Academy of Sciences, 4: 304 p.
Garman, S. (1906)
Vertebrata from the savanna of Panama. Bulletin of the Museum of Comparative Zoology at Harvard College, 46(12), 211–230
Böhlke, J.E. (1953)
A catalogue of the type specimens of Recent fishes in the Natural History Museum of Stanford University. Stanford Ichthyological Bulletin, 5(1), 1-168