MCEACHRAN, J.D. & KONSTANTINOU, H. (1996)
Survey of the variation of alar and malar thorns in skates: phylogenetic implications (Chondrichthyes: Rajoidei). Journal of Morphology, 228 (2): 165–178
DOI: 10.1002/(SICI)1097-4687(199605)228:2<165::AID-JMOR5>3.0.CO;2-4
COMPAGNO, L.J.V. (2005)
Checklist of living Chondrichthyes. In: Hamlett, W.C. (ed.) Reproductive biology and phylogeny of chondrichthyes: sharks, rays and chimaeras, Vol. 3. Endfield, USA: Science Publishers: 503–548