Menner, V.V. (1928)
Les Sélaciens du Paléogène de Manghyschlak, d'Emba et du versant oriental d'Oural. «in russian». Bulletin de la Société Impériale des Naturalistes de Moscou, Section Géologique, 6(3–4), 292–338

Zhelezko, V.I. & Kozlov, V.A. (1999)
Elasmobranhii i biostratigraphia paleogena Zauralia i Srednei Asii. (Elasmobranchii and Palaeogene biostratigraphy of Transural and Central Asia). Materialy po Stratigrafii i Paleontologii Urala, 3, 324 pp, Ekaterinburg, UrO RAN

King, C. & Iakovleva, A. & Steurbaut, E. & Heilmann-Clausen, C. & Ward, D. (2013)
The Aktulagay section, west Kazakhstan: a key site fornorthern mid-latitude Early Eocene stratigraphy. Stratigraphy, 10(3): 171–209

Malyshkina, T.P. & Ward, D.J. (2016)
The Turanian Basin in the Eocene: the new data on the fossil sharks and rays from the Kyzylkum Desert (Uzbekistan). Proceedings of the Zoological Institute, Russian Academy of Sciences, 320(1), 50–65