MERINO-RODO, D. & JANVIER, P. (1986)
Chondrichthyan and actinopterygian remains from the Lower Permian Copacabana Formation of Bolivia. Géobios, 19 (4): 479–493
DOI: 10.1016/S0016-6995(86)80005-5
JANVIER, P. (1991)
The Permian and Triassic Vertebrates of Bolivia. Revista Tecnica de Yacimientos Petroliferos Fiscales Bolivianos, 12 (3–4): 389–390
ARRATIA, G. & CIONE, A.L. (1996)
The record of fossil fishes of southern South America. Münchner Geowissenschaftliche Abhandlungen Reihe A: Geologie und Paläontologie, 30: 9–72
GINTER, M. & HAMPE, O. & DUFFIN, C.J. (2010)
Handbook of Paleoichthyology, Vol. 3D: Chondrichthyes Paleozoic Elasmobranchii: Teeth Verlag Dr. Friedrich Pfeil, pp. 168, 154 fig., 4 Tab.