Lloris, D. (1986)
Ictiofauna demersal y aspectos biogeográficos de la costa Sudoccidental de Africa (SWA/NAMIBIA). Monografías de zoología marina, Instituto de Ciencias del Mar, Barcelona, Consejo Superior de Investigaciones Científicas, 1: 429pp

Compagno, L.J.V. & Ebert, D.A. & Smale, M.J. (1989)
Guide to the sharks and rays of Southern Africa. New Holland Ltd., London 1989

Compagno, L.J.V. & Ebert, D.A. & Cowley, P.D. (1991)
Distribution of offshore demersal cartilaginous fishes (Class Chondrichthyes) of the west coast of southern Africa, with notes on their systematics. South African Journal of Marine Science, 11(1), 43–139, figs 1–24
DOI: 10.2989/025776191784287664

Stehmann, M. (1995)
First and new records of skates (Chondrichthyes, Rajiformes, Rajidae) from the West African continental slope (Morocco to South Africa), with descriptions of two new species. Archive of Fishery and Marine Research, 43(1), 1–119

Deynat, P.P. & Séret, B. (1996)
Le revêtement cutané des raies (Chondrichthyes, Elasmobranchii, Batoidea). I- Morphologie et arrangement des denticules cutanés. Annales des Sciences Naturelles – Zoologie et Biologie Animale, 17(2), 65–83

Deynat, P.P. (1998)
Le revêtement cutané des raies (Chondrichthyes, Elasmobranchii, Batoidea). II. Morphologie et arrangement des tubercules cutanés. Annales des Sciences Naturelles – Zoologie et Biologie Animale, 19(3–4), 155–172
DOI: 10.1016/S0003-4339(99)80004-5