Lloris, D. (1986)
Ictiofauna demersal y aspectos biogeográficos de la costa Sudoccidental de Africa (SWA/NAMIBIA). Monografías de zoología marina, Instituto de Ciencias del Mar, Barcelona, Consejo Superior de Investigaciones Científicas, 1: 429pp
Compagno, L.J.V. & Ebert, D.A. & Smale, M.J. (1989)
Guide to the sharks and rays of Southern Africa. New Holland Ltd., London 1989
Compagno, L.J.V. & Ebert, D.A. & Cowley, P.D. (1991)
Distribution of offshore demersal cartilaginous fishes (Class Chondrichthyes) of the west coast of southern Africa, with notes on their systematics. South African Journal of Marine Science, 11(1), 43–139, figs 1–24
DOI: 10.2989/025776191784287664
Deynat, P.P. & Séret, B. (1996)
Le revêtement cutané des raies (Chondrichthyes, Elasmobranchii, Batoidea). I- Morphologie et arrangement des denticules cutanés. Annales des Sciences Naturelles – Zoologie et Biologie Animale, 17(2), 65–83
McEachran, J.D. & Konstantinou, H. (1996)
Survey of the variation of alar and malar thorns in skates: phylogenetic implications (Chondrichthyes: Rajoidei). Journal of Morphology, 228(2), 165–178
DOI: 10.1002/(SICI)1097-4687(199605)228:2<165::AID-JMOR5>3.0.CO;2-4
Deynat, P.P. (1998)
Le revêtement cutané des raies (Chondrichthyes, Elasmobranchii, Batoidea). II. Morphologie et arrangement des tubercules cutanés. Annales des Sciences Naturelles – Zoologie et Biologie Animale, 19(3–4), 155–172
DOI: 10.1016/S0003-4339(99)80004-5