ANTUNES, M.T. & JONET, S. (1970)
Requins de l'Helvétien supérieur et du Tortonien de Lisbonne. Revista da Faculdade de Ciências de Lisboa, 16 (1): 119–280, pl. 1–20
BALBINO, A.C. (1996)
Sharks from the Middle and early Upper Miocene from Lisbon, Portugal. A check-list. Comunicações do Instituto Geológico e Mineiro, 82: 141–144
KRIWET, J. & KLUG, S. (2009)
Fossil record and origin of squaliform sharks (Chondrichthyes, Neoselachii). In: Gallucci, V.F. & McFarlane, G.A. & Bargmann, G.G. (eds). Biology and management of dogfish sharks. American Fisheries Society. Bethesda, Maryland: 19–38