Description of a new species of squaloid shark from Japan (Squalus japonicus). Proceedings of the Academy of Natural Sciences of Philadelphia, 60: 71–73
A catalogue of the fishes of the island of Formosa, or Taiwan, based on the collections of Dr. Hans Sauter. Memoirs of the Carnegie Museum, 4 (4): 159–204, Pls. 63–74
A catalogue of the fishes of Japan. Journal of the College of Science, Imperial University, Tokyo, 33: 1–497


Blainville's dogfish, Squalus blainville, from Japan, with notes on S. mitsukurii and S. japonicus. Japanese Journal of Ichthyology, 26 (1): 26–42

Some aspects of reproduction in the pointed-snout dogfish Squalus japonicus taken of Nagasaki and Choshi. Bulletin of the Japanese Society of Scientific Fisheries, 47 (9): 1157–1164

Preliminary investigation of sharks and rays in Southeast Asia - Philippines. (in japanese) Report of Japanese Society for Elasmobranch Studies, 15: 7–11, figs 1–6


FAO Species Catalogue. Vol 4: Sharks of the world, Part 1 - Hexanchiformes to Lamniformes. FAO Fisheries Synopsis No. 125, 4 (1): 1–250

Shark myoglobins-I. Isolation and characterization of myoglobins from the sharks, Squalus japonicus and Proscyllium habereri. Comparative Biochemistry and Physiology – Part B, Comparative Biochemistry, 78 (1): 163–166
DOI: 10.1016/0305-0491(84)90161-5

Sharks, Skates, Rays and Chimaeras Taken from Continental Slopes around Japan. (in japanese) Report of Japanese Society for Elasmobranch Studies, 19: 1–8


Dogfishes Squalus (Squalidae), of the Atlantic Ocean and comparative notes on the species of this genus from other regions. Journal of Ichthyology, 27 (1): 1–18


A key to the species of the dogfishes (Squaloidei) from Japanese waters. (in japanese) Report of Japanese Society for Elasmobranch Studies, 26: 1–35
Interrelationships of the Etmopteridae (Chondrichthyes, Squaliformes). In: Pratt, H.L., Gruber, S.H., Taniuchi, T., editors. Elasmobranchs as living resources: advances in the biology, ecology, systematics, and the status of the fisheries: NOAA Technical Report NMFS, 90: 347–356

Fauna and Zoogeography of Deep-Benthic chondrichthyan Fishes around the Japanese Archipelago. Japanese Journal of Ichthyology, 39 (1): 37–48
DOI: 10.1007/BF02905632

Squalean phylogeny: a new framework of "squaloid" sharks and related taxa. Hokkaido University Press, 151 pp.

Deep-sea chondrichthyans collected from the waters around the Okinawa Islands: results of catch analysis of bottom longlines. Bulletin of the Seikai National Fisheries Research Institute, 71: 51–65
Molecular Phylogeny of the Sharks and Rays of Superorder Squalea Based on Mitochondrial Cytochrome b Gene. Fisheries Science, 62 (3): 340–343
DOI: 10.2331/fishsci.62.340

Classification of the recent Elasmobranchii. Copyright Brian Mould 1997

Annotated checklist of deep-water fishes from Suruga Bay, Japan. National Science Museum Monographs, 12: 269–318, pls 1–2
FAO species identification guide for fishery purposes. The living marine resources of the Western Central Pacific. Volume 2. Cephalopods, crustaceans, holothurians and sharks. Rome, FAO. 1998: 687–1396

Part Sharks: Hexanchidae, Echinorhinidae, Squalidae, Squatinidae, Heterodontidae, Parascyllidae, Bracheluridae, Orectolobidae, Hemiscyllidae, Odontaspididae, Scyliorhinidae, Proscyllidae, Triakidae, In: K.E. Carpenter and V.H. Niem (eds.), FAO Identification Guide for Fishery Purposes. The Living Marine Resources of the Western Central Pacific. Food and Agriculture Organization, Rome: 1195–1232, 1235–1259, 1264–1267, 1279–1295, 1297–1304, 1312–1360

Chondrichthyans of the Ryukyu Islands, Japan. In: Seret, B. and Sire, J.Y. (eds) Indo-Pacific fish biology: Proceedings of the 5th Indo-Pacific Fisheries Conference, Noumea, 1997. Ichthyological Society of France, Paris: 351–365

Life history patterns and correlations in sharks. Reviews in Fisheries Science, 8: 299–344
DOI: 10.1080/10408340308951115
A checklist of the fishes of the South China Sea. Raffles Bulletin of Zoology, Supplement 8: 569–667

A catalogue of fishes from the East China and Yellow seas with their local names. Seikai National Fisheries Research Institute , Fisheries Research Agency, Nagasaki, iii+42 pp. (in Japanese)

Checklist of living Chondrichthyes. In: Hamlett, W.C. (ed.) Reproductive biology and phylogeny of chondrichthyes: sharks, rays and chimaeras, Vol. 3. Endfield, USA: Science Publishers: 503–548
A Collation and summarization of available data on deepwater Chondrichthyans: Biodiversity, life history and fisheries. Biodiversity, Life History and Fisheries IUCN SSC Shark Specialist Group for the Marine Conservation Biology Institute, 1–137.

DNA barcoding Australasian chondrichthyans: results and potential uses in conservation. Marine and Freshwater Research, 59 (1): 57–71
DOI: 10.1071/MF07148
Squalus japonicus In: IUCN 2012. IUCN Red List of Threatened Species. Version 2012.2. <www.iucnredlist.org>

Shark tales: A molecular species-level phylogeny of sharks (Selachimorpha, Chondrichthyes). Molecular Phylogenetics and Evolution, 58 (2): 207–217
DOI: 10.1016/j.ympev.2010.11.018
A DNA sequence based approach to the identification of shark and ray species and its implications for global elasmobranch diversity and parasitology. Bulletin of the American Museum of Natural History, 367: 262 pp., 102 figures, 5 tables
An illustrated list of the Kagoshima University Museum specimens of Chondrichthyes collected from Kagoshima Prefecture, southern Japan. [In Japanese.] Nature of Kagoshima, 38: 119–138

Sharks of the World – A fully illustrated guide. Wild Nature Press, ISBN 978–0–9573946–0–5: 528pp
An annotated checklist of the chondrichthyans of Taiwan. Zootaxa, 3752: 279–386
DOI: 10.11646/zootaxa.3752.1.17

Fishes of Japan with Pictorial Keys to the Species. Third Edition. Tokyo: Tokai University Press. 2428 pp.
A DNA sequence-based identification checklist for Taiwanese chondrichthyans. Zootaxa, 3752: 256–278
DOI: 10.11646/zootaxa.3752.1.16

The effect of habitat on modern shark diversification. Journal of Evolutionary Biology, 27 (8): 1536–1548
DOI: 10.1111/jeb.12405
Global pattern of phylogenetic species composition of shark and its conservation priority. Ecology and Evolution, 5 (19): 4455–4465
DOI: 10.1002/ece3.1724
The ichthyofauna of the Uji Islands, East China Sea: 148 new records of fishes with notes on biogeographical implications. Memoirs of the Faculty of Fisheries, Kagoshima University, 64: 10–34

Ultrasound and physical models shed light on the respiratory system of embryonic dogfishes. Zoology (Jena), 119 (1): 36–41
DOI: 10.1016/j.zool.2015.09.002

Annotated checklist of the living sharks, batoids and chimaeras (Chondrichthyes) of the world, with a focus on biogeographical diversity. Journal of Fish Biology, 88 (3): 837–1037
DOI: 10.1111/jfb.12874

Molecular diversity and distribution of eastern Atlantic and Mediterranean dogfishes Squalus highlight taxonomic issues in the genus. Zoologica Scripta, 46 (4): 414-428
DOI: 10.1111/zsc.12224

Sustainability of threatened species displayed in public aquaria, with a case study of Australian sharks and rays. Reviews in Fish Biology and Fisheries, 28 (1): 137–151
DOI: 10.1007/s11160-017-9501-2

An identification key to elasmobranch genera based on dental morphological characters Part A: Squalomorph sharks (Superorder Squalomorphii). Bulletin of Fish Biology, 18 (1/2): 77-105

Species composition and biomass density of mesopelagic nekton of the South China Sea continental slope. Deep Sea Research Part II: Topical Studies in Oceanography, 167: 105-120
DOI: 10.1016/j.dsr2.2018.06.008
Rapid detection of CITES-listed shark fin species by loop-mediated isothermal amplification assay with potential for field use. Scientific Reports, 10 (1): 4455
DOI: 10.1038/s41598-020-61150-8
Distinct Responses of Elasmobranchs and Ray-Finned Fishes to Long-Term Global Change. Frontiers in Ecololgy and Evolution, 7: 513
DOI: 10.3389/fevo.2019.00513
First Record of the Rare Parasitic Isopod Elthusa splendida (Cymothoidae) from the Pacific Ocean, Based on a Specimen Found in a Museum Shark Collection. Species Diversity, 25 (2): 343
DOI: 10.12782/specdiv.25.343

Body forms in sharks (Chondrichthyes: Elasmobranchii) and their functional, ecological, and evolutionary implications. Zoology, 140: 125799
DOI: 10.1016/j.zool.2020.125799
Squalus shiraii sp. nov. (Squaliformes, Squalidae), a new species of dogfish shark from Japan with regional nominal species revisited. Zoosystematics and Evolution, 96 (2): 275-311
DOI: 10.3897/zse.96.51962

Filling gaps in the biology and habitat use of two spurdog sharks (Squalus japonicus and Squalus brevirostris) in the East China Sea. Marine and Freshwater Research, 71 (12): 1719-1731
DOI: 10.1071/MF19131

Re-evaluation of deep-sea dogfishes (genus Squalus) in Japan using phylogenetic inferenc. Deep Sea Research Part I: Oceanographic Research Papers, in press
DOI: 10.1016/j.dsr.2020.103261