Malyshkina, T.P. & Ward, D.J. (2016)
The Turanian Basin in the Eocene: the new data on the fossil sharks and rays from the Kyzylkum Desert (Uzbekistan). Proceedings of the Zoological Institute, Russian Academy of Sciences, 320(1), 50–65

King, C. & Iakovleva, A. & Steurbaut, E. & Heilmann-Clausen, C. & Ward, D. (2013)
The Aktulagay section, west Kazakhstan: a key site fornorthern mid-latitude Early Eocene stratigraphy. Stratigraphy, 10(3): 171–209

Zhelezko, V.I. & Kozlov, V.A. (1999)
Elasmobranhii i biostratigraphia paleogena Zauralia i Srednei Asii. (Elasmobranchii and Palaeogene biostratigraphy of Transural and Central Asia). Materialy po Stratigrafii i Paleontologii Urala, 3, 324 pp, Ekaterinburg, UrO RAN

Menner, V.V. (1928)
Les Sélaciens du Paléogène de Manghyschlak, d'Emba et du versant oriental d'Oural. «in russian». Bulletin de la Société Impériale des Naturalistes de Moscou, Section Géologique, 6(3–4), 292–338