Guinot, G. & Amoudji, Y.Z. & Kassegne, K.E. & Sarr, R. & Johnson, A.K.C. & da Costa, P.Y.D. & Hautier, L. (2020)
Elasmobranchs from the upper Palaeocene of Togo. Geobios, 58, 17–37
DOI: 10.1016/j.geobios.2019.12.003

Underwood, C.J. & Ward, D.J. & King, C. & Antar, S.M. & Zalmout, I.S. & Gingerich, P.D. (2011)
Shark and ray faunas in the Middle and Late Eocene of the Fayum Area, Egypt. Proceedings of the Geologists' Association, 122(1), 47–66
DOI: 10.1016/j.pgeola.2010.09.004

Antar, M.S.M. (2011)
Paleo-environments of the exposed Eocene Sediments between Wadi El-Hitan and east Siwa in the Egyptian Western Desert based on their faunal content especially the vertebrates. Thesis, Zagazig University, Faculty of Science, Geology Department