Orlov, A.M. & Cotton, C.F. & Shevernitsky, D.A. (2011)
Sexual dimorphism of external morphological characters in some deepwater skates (Rajidae, Rajiformes, Chondrichthyes) of the North Atlantic.
Moscow University Biological Sciences Bulletin, 65(1), 40–44 DOI: 10.3103/S0096392510010086