da Cruz, V.P. & Shimabukuro-Dias, C.K. & Oliveira, C. & Foresti, F. (2011)
Karyotype description and evidence of multiple sex chromosome system X1X1X2X2/X1X2Y in Potamotrygon aff. motoro and P. falkneri (Chondrichthyes: Potamotrygonidae) in the upper Parana River basin, Brazil.
Neotropical Ichthyology, 9(1), 201–208