Towner, A. & Micarelli, P. & Hurwitz, D. & Smale, M.J. & Booth, A.J. & Stopforth, C. & Jacobs, E. & Reinero, F.R. & Ricci, V. & Di Bari, A. & Gavazzi, S. & Carugno, G. & Mahrer, M. & Gennari, E. (2024)
Further insights into killer whales Orcinus orca preying on white sharks Carcharodon carcharias in South Africa. African Journal of Marine Science, in press
DOI: 10.2989/1814232X.2024.2311272

Soekoe, M. & Smale, M.J. & Potts, W.M. (2022)
Highly conserved tooth morphology in allopatric elasmobranch populations despite contrasting diets—a case of Triakis megalopterus in southern Africa. Environmental Biology of Fishes, 105(7), 821–850
DOI: 10.1007/s10641-022-01288-8

Towner, A.V. & Watson, R.G.A. & Kock, A.A. & Papastamatiou, Y. & Sturup, M. & Gennari, E. & Baker, K. & Booth, T. & Dicken, M. & Chivell, W. & Elwen, S. & Kaschke, T. & Edwards, D. & Smale, M.J. (2022)
Fear at the top: killer whale predation drives white shark absence at South Africa's largest aggregation site. African Journal of Marine Science, 44(2), 139–152
DOI: 10.2989/1814232x.2022.2066723

Kock, A.A. & Lombard, A.T. & Daly, R. & Goodall, V. & Meyer, M. & Johnson, R. & Fischer, C. & Koen, P. & Irion, D. & Gennari, E. & Towner, A. & Jewell, O.J.D. & da Silva, C. & Dicken, M.L. & Smale, M.J. & Photopoulou, T. (2022)
Sex and Size Influence the Spatiotemporal Distribution of White Sharks, With Implications for Interactions With Fisheries and Spatial Management in the Southwest Indian Ocean. Frontiers in Marine Science, 9, Article 811985
DOI: 10.3389/fmars.2022.811985

Queiroz, N. & Humphries, N.E. & Couto, A. & Vedor, M. & da Costa, I. & Sequeira, A.M.M. & Mucientes, G. & Santos, A.M. & Abascal, F.J. & Abercrombie, D.L. & Abrantes, K. & Acuna-Marrero, D. & Afonso, A.S. & Afonso, P. & Anders, D. & Araujo, G. & Arauz, R. & Bach, P. & Barnett, A. & Bernal, D. & Berumen, M.L. & Lion, S.B. & Bezerra, N.P.A. & Blaison, A.V. & Block, B.A. & Bond, M.E. & Bonfil, R. & Bradford, R.W. & Braun, C.D. & Brooks, E.J. & Brooks, A. & Brown, J. & Bruce, B.D. & Byrne, M.E. & Campana, S.E. & Carlisle, A.B. & Chapman, D.D. & Chapple, T.K. & Chisholm, J. & Clarke, C.R. & Clua, E.G. & Cochran, J.E.M. & Crochelet, E.C. & Dagorn, L. & Daly, R. & Cortes, D.D. & Doyle, T.K. & Drew, M. & Duffy, C.A.J. & Erikson, T. & Espinoza, E. & Ferreira, L.C. & Ferretti, F. & Filmalter, J.D. & Fischer, G.C. & Fitzpatrick, R. & Fontes, J. & Forget, F. & Fowler, M. & Francis, M.P. & Gallagher, A.J. & Gennari, E. & Goldsworthy, S.D. & Gollock, M.J. & Green, J.R. & Gustafson, J.A. & Guttridge, T.L. & Guzman, H.M. & Hammerschlag, N. & Harman, L. & Hazin, F.H.V. & Heard, M. & Hearn, A.R. & Holdsworth, J.C. & Holmes, B.J. & Howey, L.A. & Hoyos, M. & Hueter, R.E. & Hussey, N.E. & Huveneers, C. & Irion, D.T. & Jacoby, D.M.P. & Jewell, O.J.D. & Johnson, R. & Jordan, L.K.B. & Joyce, W. & Daly, C.A.K. & Ketchum, J.T. & Klimley, A.P. & Kock, A.A. & Koen, P. & Ladino, F. & Lana, F.O. & Lea, J.S.E. & Llewellyn, F. & Lyon, W.S. & MacDonnell, A. & Macena, B.C.L. & Marshall, H. & McAllister, J.D. & Meyer, M.A. & Morris, J.J. & Nelson, E.R. & Papastamatiou, Y.P. & Penaherrera-Palma, C. & Pierce, S.J. & Poisson, F. & Quintero, L.M. & Richardson, A.J. & Rogers, P.J. & Rohner, C.A. & Rowat, D.R.L. & Samoilys, M. & Semmens, J.M. & Sheaves, M. & Shillinger, G. & Shivji, M. & Singh, S. & Skomal, G.B. & Smale, M.J. & Snyders, L.B. & Soler, G. & Soria, M. & Stehfest, K.M. & Thorrold, S.R. & Tolotti, M.T. & Towner, A. & Travassos, P. & Tyminski, J.P. & Vandeperre, F. & Vaudo, J.J. & Watanabe, Y.Y. & Weber, S.B. & Wetherbee, B.M. & White, T.D. & Williams, S. & Zarate, P.M. & Harcourt, R. & Hays, G.C. & Meekan, M.G. & Thums, M. & Irigoien, X. & Eguiluz, V.M. & Duarte, C.M. & Sousa, L.L. & Simpson, S.J. & Southall, E.J. & Sims, D.W. (2021)
Reply to: Shark mortality cannot be assessed by fishery overlap alone. Nature, 595(7866), E8–E16
DOI: 10.1038/s41586-021-03397-3

da Silva, C. & Attwood, C.G. & Wintner, S.P. & Wilke, C.G. & Winker, H. & Smale, M.J. & Kerwath, S.E. (2021)
Life history of Mustelus mustelus in the Langebaan Lagoon marine protected area. Marine and Freshwater Research, 72(8), 1142–1159
DOI: 10.1071/mf20277

Queiroz, N. & Humphries, N.E. & Couto, A. & Vedor, M. & Costa, I. & Sequeira, A.M.M. & Mucientes, G. & Santos, A.M. & Abascal, F.J. & Abercrombie, D.L. & Abrantes, K. & Acuna-Marrero, D. & Afonso, A.S. & Afonso, P. & Anders, D. & Araujo, G. & Arauz, R. & Bach, P. & Barnett, A. & Bernal, D. & Berumen, M.L. & Lion, S.B. & Bezerra, N.P.A. & Blaison, A.V. & Block, B.A. & Bond, M.E. & Bonfil, R. & Bradford, R.W. & Braun, C.D. & Brooks, E.J. & Brooks, A. & Brown, J. & Bruce, B.D. & Byrne, M.E. & Campana, S.E. & Carlisle, A.B. & Chapman, D.D. & Chapple, T.K. & Chisholm, J. & Clarke, C.R. & Clua, E.G. & Cochran, J.E.M. & Crochelet, E.C. & Dagorn, L. & Daly, R. & Cortes, D.D. & Doyle, T.K. & Drew, M. & Duffy, C.A.J. & Erikson, T. & Espinoza, E. & Ferreira, L.C. & Ferretti, F. & Filmalter, J.D. & Fischer, G.C. & Fitzpatrick, R. & Fontes, J. & Forget, F. & Fowler, M. & Francis, M.P. & Gallagher, A.J. & Gennari, E. & Goldsworthy, S.D. & Gollock, M.J. & Green, J.R. & Gustafson, J.A. & Guttridge, T.L. & Guzman, H.M. & Hammerschlag, N. & Harman, L. & Hazin, F.H.V. & Heard, M. & Hearn, A.R. & Holdsworth, J.C. & Holmes, B.J. & Howey, L.A. & Hoyos, M. & Hueter, R.E. & Hussey, N.E. & Huveneers, C. & Irion, D.T. & Jacoby, D.M.P. & Jewell, O.J.D. & Johnson, R. & Jordan, L.K.B. & Jorgensen, S.J. & Joyce, W. & Daly, C.A.K. & Ketchum, J.T. & Klimley, A.P. & Kock, A.A. & Koen, P. & Ladino, F. & Lana, F.O. & Lea, J.S.E. & Llewellyn, F. & Lyon, W.S. & MacDonnell, A. & Macena, B.C.L. & Marshall, H. & Mcallister, J.D. & McAuley, R. & Meyer, M.A. & Morris, J.J. & Nelson, E.R. & Papastamatiou, Y.P. & Patterson, T.A. & Penaherrera-Palma, C. & Pepperell, J.G. & Pierce, S.J. & Poisson, F. & Quintero, L.M. & Richardson, A.J. & Rogers, P.J. & Rohner, C.A. & Rowat, D.R.L. & Samoilys, M. & Semmens, J.M. & Sheaves, M. & Shillinger, G. & Shivji, M. & Singh, S. & Skomal, G.B. & Smale, M.J. & Snyders, L.B. & Soler, G. & Soria, M. & Stehfest, K.M. & Stevens, J.D. & Thorrold, S.R. & Tolotti, M.T. & Towner, A. & Travassos, P. & Tyminski, J.P. & Vandeperre, F. & Vaudo, J.J. & Watanabe, Y.Y. & Weber, S.B. & Wetherbee, B.M. & White, T.D. & Williams, S. & Zarate, P.M. & Harcourt, R. & Hays, G.C. & Meekan, M.G. & Thums, M. & Irigoien, X. & Eguiluz, V.M. & Duarte, C.M. & Sousa, L.L. & Simpson, S.J. & Southall, E.J. & Sims, D.W. (2019)
Global spatial risk assessment of sharks under the footprint of fisheries. Nature, 572(7770), 461
DOI: 10.1038/s41586-019-1444-4

Gennari, E. & Kock, A.A. & Smale, M.J. & Towner, A. & Khan, N. & Bester, L.A. & Johnson, R. & Fischer, C. & Meyer, M. & Morse, P. (2019)
Antibiotic sensitivity of bacteria isolated from the oral cavities of live white sharks (Carcharodon carcharias) in South African waters. South African Journal of Science, 115(11/12), 5972
DOI: 10.17159/sajs.2019/5972

Daly, R. & Smale, M.J. & Singh, S. & Anders, D. & Shivji, M. & Daly, C.A.K. & Lea, J.S.E. & Sousa, L.L. & Wetherbee, B.M. & Fitzpatrick, R. & Clarke, C.R. & Sheaves, M. & Barnett, A. (2018)
Refuges and risks: Evaluating the benefits of an expanded MPA network for mobile apex predators. Diversity and Distributions, 24(9), 1217–1230
DOI: 10.1111/ddi.12758

Dicken, M.L. & Winker, H. & Smale, M.J. & Cliff, G. (2018)
Sharks caught in the KwaZulu-Natal bather protection programme, South Africa. 14. The smooth hammerhead shark Sphyrna zygaena (Linnaeus). African Journal of Marine Science, 40(2), 157–174
DOI: 10.2989/1814232x.2018.1470031

Huveneers, C. & Apps, K. & Becerril-García, E.E. & Bruce, B. & Butcher, P.A. & Carlisle, A.B. & Chapple, T.K. & Christiansen, H.M. & Cliff, G. & Curtis, T.H. & Daly-Engel, T.S. & Dewar, H. & Dicken, M.L. & Domeier, M.L. & Duffy, C.A.J. & Ford, R. & Francis, M.P. & French, G.C.A. & Galván-Magaña, F. & García-Rodríguez, E. & Gennari, E. & Graham, B. & Hayden, B. & Hoyos-Padilla, E.M. & Hussey, N.E. & Jewell, O.J.D. & Jorgensen, S.J. & Kock, A.A. & Lowe, C.G. & Lyons, K. & Meyer, L. & Oelofse, G. & Oñate-González, E.C. & Oosthuizen, H. & O'Sullivan, J.B. & Ramm, K. & Skomal, G. & Sloan, S. & Smale, M.J. & Sosa-Nishizaki, O. & Sperone, E. & Tamburin, E. & Towner, Av. & Wcisel, M.A. & Weng, K.C. & Werry, J.M. (2018)
Future Research Directions on the “Elusive” White Shark. Frontiers in Marine Science, 5, Article 455
DOI: 10.3389/fmars.2018.00455

Filmalter, J.D. & Cowley, P.D. & Potier, M. & Ménard, F. & Smale, M.J. & Cherel, Y. & Dagorn, L. (2017)
Feeding ecology of silky sharks Carcharhinus falciformis associated with floating objects in the western Indian Ocean. Journal of Fish Biology, 90(4), 1321–1337
DOI: 10.1111/jfb.13241

Dicken, M.L. & Hussey, N.E. & Christiansen, H.M. & Smale, M.J. & Nkabi, N. & Cliff, G. & Wintner, S.P. (2017)
Diet and trophic ecology of the tiger shark (Galeocerdo cuvier) from South African waters. PLoS ONE, 12(6), Article e0177897
DOI: 10.1371/journal.pone.0177897

Towner, A.V. & Leos-Barajas, V. & Langrock, R. & Schick, R.S. & Smale, M.J. & Kaschke, T. & Jewell, O.J.D. & Papastamatiou, Y.P. (2016)
Sex-specific and individual preferences for hunting strategies in white sharks. Functional Ecology, 30, 1397–1407
DOI: 10.1111/1365-2435.12613

Smale, M.J. & Dicken, M.L. & Booth, A.J. (2015)
Seasonality, behaviour and philopatry of spotted ragged‑tooth sharks Carcharias taurus in Eastern Cape nursery areas, South Africa. African Journal of Marine Science, 37(2), 219–231
DOI: 10.2989/1814232X.2015.1043342

Daly, R. & Froneman, P.W. & Cowley, P.D. & Smale, M.J. (2014)
Niche width and habitat use of adult bull sharks, Carcharhinus leucas, on the east coast of southern Africa [Abstract]. In Programm and Abstracts of Shark International, Durban 2014: 43

Dicken, M.L. & Smale, M.J. & Bradshaw, M. (2014)
An investigation into the occurrence and distribution of white sharks, Carcharodon carcharias, in Algoa Bay, South Africa [Abstract]. In Programm and Abstracts of Shark International, Durban 2014: 51

Smale, M.J. & Dicken, M.L. & Booth, A.J. (2014)
Seasonality, behavior and philopatry of raggedtooth sharks, Carcharias taurus in Eastern Cape nursery areas, South Africa [Abstract]. In Programm and Abstracts of Shark International, Durban 2014: 163

Soekoe, M. & Potts, W.M. & Smale, M.J. (2014)
When science and architecture meet: the development of a truss network for sharks [Abstract]. In Programm and Abstracts of Shark International, Durban 2014: 167

Towner, A.V. & Underhill, L. & Smale, M.J. (2014)
Great white sharks, Carcharodon carcharias, in Gansbaai, South Africa: Environmental influences and changes over time [Abstract]. In Programm and Abstracts of Shark International, Durban 2014: 174

Daly, R. & Smale, M.J. & Cowley, P.D. & Froneman, P.W. (2014)
Residency Patterns and Migration Dynamics of Adult Bull Sharks (Carcharhinus leucas) on the East Coast of Southern Africa. PLoS ONE, 9(10), Article e109357
DOI: 10.1371/journal.pone.0109357

Daly, R. & Smale, M.J. (2013)
Evaluation of an underwater biopsy probe for collecting tissue samples from bull sharks Carcharhinus leucas. African Journal of Marine Science, 35(1), 129–132
DOI: 10.2989/1814232X.2013.769910

Towner, A.V. & Underhill, L.G. & Jewell, O.J.D. & Smale, M.J. (2013)
Environmental Influences on the Abundance and Sexual Composition of White Sharks Carcharodon carcharias in Gansbaai, South Africa. PLoS ONE, 8(8), Article e71197
DOI: 10.1371/journal.pone.0071197

Dicken, M.L. & Smale, M.J. & Booth, A.J. (2013)
White sharks Carcharodon carcharias at Bird Island, Algoa Bay, South Africa. African Journal of Marine Science, 35(2), 175–182
DOI: 10.2989/1814232X.2013.800579

Daly, R. & Froneman, P.W. & Smale, M.J. (2013)
Comparative Feeding Ecology of Bull Sharks (Carcharhinus leucas) in the Coastal Waters of the Southwest Indian Ocean Inferred from Stable Isotope Analysis. PLoS ONE, 8(10), Article e78229
DOI: 10.1371/journal.pone.0078229

Dicken, M.L. & Bradshaw, M. & Smale, M.J. (2013)
White shark (Carcharodon carcharias)-inflicted bite wounds observed on Cape fur seals (Arctocephalus pusillus pusillus) at Black Rocks, Algoa Bay, South Africa. African Zoology, 48(2), 418–426
DOI: 10.3377/004.048.0203

Smale, M.J. & Booth, A.J. & Farquhar, M.R. & Meyer, M.R. & Rochat, L. (2012)
Migration and habitat use of formerly captive and wild raggedtooth sharks (Carcharias taurus) on the southeast coast of South Africa. Marine Biology Research, 8(2), 115–128
DOI: 10.1080/17451000.2011.617756

Dicken, M.L. & Smale, M.J. & Booth, A.J. (2012)
Long-term catch and effort trends in Eastern Cape Angling Week competitions. African Journal of Marine Science, 34(2), 259–268
DOI: 10.2989/1814232X.2012.709961

Smale, M.J. & Cliff, G. (2012)
Chapter 4: White Sharks and Cephalopod Prey: Indicators of Habitat Use? In M.L. Domeier (Ed.), Global Perspectives on the Biology and Life History of the White Shark (pp. 51–58). CRC Press

Towner, A. & Smale, M.J. & Jewell, O. (2012)
Chapter 6: Boat-Strike Wound Healing in Carcharodon carcharias. In M.L. Domeier (Ed.), Global Perspectives on the Biology and Life History of the White Shark (pp. 77–84). CRC Press

Booth, A.J. & Foulis, A.J. & Smale, M.J. (2011)
Age validation, Size, mortality, and demographic modeling of spotted gully shark (Triakis megalopterus) from the southeast coast of South Africa. Fishery Bulletin, 109(1), 101–112

Dicken, M.L. & Nance, S.P. & Smale, M.J. (2011)
Sessile biofouling on tags from recaptured raggedtooth sharks (Carcharias taurus) and their effects on tagging studies. Marine and Freshwater Research, 62(4), 359–364
DOI: 10.1071/MF10255

Peddemors, V.M. & Abernathy, K. & Macdougal, K. & Marshall, G.J. & Masens, O.J. & Smale, M.J. & Smoothey, A.F. & Gray, C.A. (2011)
Animal-borne cameras: An opportunity to investigate detailed behaviour of sharks [Abstract]. In Program and Abstracts for the 2011 Workshop and Conference of the Oceania Chondrichthyan Society, 13th–15th September 2011, Sea World Resort and Water Park, Gold Coast, Queensland, Australia.

Smale, M.J. (2009)
Amblyraja robertsi IUCN 2012, IUCN Red List of Threatened Species, Version 2012.2, https://www.iucnredlist.org

Smale, M.J. (2009)
Bathyraja eatonii IUCN 2012, IUCN Red List of Threatened Species, Version 2012.2, https://www.iucnredlist.org

Smale, M.J. (2009)
Bathyraja irrasa IUCN 2012, IUCN Red List of Threatened Species, Version 2012.2, https://www.iucnredlist.org

Smale, M.J. & Holtzhausen, J.A. & Mccormack, C. (2009)
Bathyraja smithii IUCN 2012, IUCN Red List of Threatened Species, Version 2012.2, https://www.iucnredlist.org

Smale, M.J. (2009)
Carcharhinus amblyrhynchos IUCN 2012, IUCN Red List of Threatened Species, Version 2012.2, https://www.iucnredlist.org

Smale, M.J. & Pheeha, S. (2009)
Cruriraja durbanensis IUCN 2012, IUCN Red List of Threatened Species, Version 2012.2, https://www.iucnredlist.org

Smale, M.J. (2009)
Cruriraja parcomaculata IUCN 2012, IUCN Red List of Threatened Species, Version 2012.2, https://www.iucnredlist.org

Smale, M.J. (2009)
Cruriraja triangularis IUCN 2012, IUCN Red List of Threatened Species, Version 2012.2, https://www.iucnredlist.org

Smale, M.J. (2009)
Dasyatis chrysonota IUCN 2012, IUCN Red List of Threatened Species, Version 2012.2, https://www.iucnredlist.org

Smale, M.J. (2009)
Dipturus doutrei IUCN 2012, IUCN Red List of Threatened Species, Version 2012.2, https://www.iucnredlist.org

Smale, M.J. (2009)
Dipturus springeri IUCN 2012, IUCN Red List of Threatened Species, Version 2012.2, https://www.iucnredlist.org

Smale, M.J. & Compagno, L.J.V. & Mccormack, C. (2009)
Leucoraja compagnoi IUCN 2012, IUCN Red List of Threatened Species, Version 2012.2, https://www.iucnredlist.org

Smale, M.J. (2009)
Leucoraja wallacei IUCN 2012, IUCN Red List of Threatened Species, Version 2012.2, https://www.iucnredlist.org

Holtzhausen, H.A. & Smale, M.J. (2009)
Narke capensis IUCN 2012, IUCN Red List of Threatened Species, Version 2012.2, https://www.iucnredlist.org

Smale, M.J. & Ungaro, N. & Serena, F. & Dulvy, N.K. & Tinti, F. & Bertozzi, M. & Mancusi, C. & Notarbartolo Di Sciara, G. (2009)
Raja miraletus IUCN 2012, IUCN Red List of Threatened Species, Version 2012.2, https://www.iucnredlist.org

Smale, M.J. (2009)
Raja straeleni IUCN 2012, IUCN Red List of Threatened Species, Version 2012.2, https://www.iucnredlist.org

Notarbartolo Di Sciara, G. & Serena, F. & Ungaro, N. & Ferretti, F. & Holtzhausen, H.A. & Smale, M.J. (2009)
Torpedo nobiliana IUCN 2012, IUCN Red List of Threatened Species, Version 2012.2, https://www.iucnredlist.org

Dicken, M.L. & Booth, A.J. & Smale, M.J. (2009)
Factors affecting recapture rates of raggedtooth sharks Carcharias taurus tagged off the east coast of South Africa. African Journal of Marine Science, 31(3), 365–372
DOI: 10.2989/AJMS.2009.31.3.9.997

Dicken, M.L. & Booth, A.J. & Smale, M.J. (2008)
Estimates of juvenile and adult raggedtooth shark (Carcharias taurus) abundance along the east coast of South Africa. Canadian Journal of Fisheries and Aquatic Sciences, 65(4), 621–632
DOI: 10.1139/f07-190

Smale, M.J. (2008)
Pelagic Shark Fisheries in the Indian Ocean. In M.D. Camhi, E.K. Pikitch, E.A. Babcock (Eds.) Sharks of the Open Ocean: Biology, Fisheries and Conservation. (pp. 247–259). Blackwell Publishing, Oxford, U.K.

Dicken, M.L. & Booth, A.J. & Smale, M.J. & Cliff, G. (2007)
Spatial and seasonal distribution patterns of juvenile and adult raggedtooth sharks (Carcharias taurus) tagged off the east coast of South Africa. Marine and Freshwater Research, 58(1), 127–134
DOI: 10.1071/MF06018

Smale, M.J. & Kulka, D.W. (2007)
Malacoraja spinacidermis IUCN 2012, IUCN Red List of Threatened Species, Version 2012.2, https://www.iucnredlist.org

Dicken, M.L. & Booth, A.J. & Smale, M.J. (2006)
Preliminary observations of tag shedding, tag reporting, tag wounds, and tag biofouling for raggedtooth sharks (Carcharias taurus) tagged off the east coast of ... shark (Carcharias taurus), tagged off the east coast of South Africa. ICES Journal of Marine Science, 63, 1640–1648
DOI: 10.1016/j.icesjms.2006.06.009

Baum, J.K. & Medina, E. & Musick, J.A. & Smale, M.J. (2006)
Carcharhinus longimanus IUCN 2012, IUCN Red List of Threatened Species, Version 2012.2, https://www.iucnredlist.org

Smale, M.J. (2006)
Mustelus palumbes IUCN 2012, IUCN Red List of Threatened Species, Version 2012.2, https://www.iucnredlist.org

Burgess, G.H. & Holtzhausen, H.A. & Smale, M.J. (2006)
Rhinobatos annulatus IUCN 2012, IUCN Red List of Threatened Species, Version 2012.2, https://www.iucnredlist.org

Smale, M.J. (2006)
Torpedo sinuspersici IUCN 2012, IUCN Red List of Threatened Species, Version 2012.2, https://www.iucnredlist.org

Dicken, M.L. & Smale, M.J. & Booth, A.J. (2006)
Shark fishing effort and catch of the ragged-tooth shark Carcharias taurus in the South African competitive shore-angling fishery. African Journal of Marine Science, 28(3–4), 589–601
DOI: 10.2989/18142320609504209

Dicken, M.L. & Smale, M.J. & Booth, A.J. (2006)
Spatial and seasonal distribution patterns of the ragged-tooth shark Carcharias taurus along the coast of South Africa. African Journal of Marine Science, 28(3–4), 603–616
DOI: 10.2989/18142320609504210

Smale, M.J. (2005)
Species accounts: Carcharhinus amblyrhynchos, Carcharhinus longimanus, Triaenodon obesus In Fowler, S.L., Cavanagh, R.D., Camhi, M., Burgess, G.H., Cailliet, G.M., Fordham, S.V., Simpfendorfer, C.A. and Musick, J.A.(comp. and Ed.). Sharks, Rays and Chimaeras: The Status of the Chondrichthyan Fishes. Status Survey. IUCN SSC Shark Specialist Group. IUCN, Gland, Switzerland and Cambridge, UK: 284–285, 295–296, 314

Smale, M.J. (2005)
Triaenodon obesus IUCN 2012, IUCN Red List of Threatened Species, Version 2012.2, https://www.iucnredlist.org

Smale, M.J. (2005)
The diet of the ragged-tooth shark Carcharias taurus Rafinesque 1810 in the Eastern Cape, South Africa. African Journal of Marine Science, 27(1), 331–336
DOI: 10.2989/18142320509504091

de Bruyn, P. & Dudley, S.F.J. & Cliff, G. & Smale, M.J. (2005)
Sharks caught in the protective gill nets off KwaZulu-Natal, South Africa. 11. The scalloped hammerhead shark Sphyrna lewini (Griffith and Smith). African Journal of Marine Science, 27(3), 517–528
DOI: 10.2989/18142320509504112

Dudley, S.F.J. & Cliff, G. & Zungu, M.P. & Smale, M.J. (2005)
Sharks caught in the protective gill nets off KwaZulu-Natal, South Africa. 10. The dusky shark Carcharhinus obscurus (Lesueur 1818). African Journal of Marine Science, 27(1), 107–128
DOI: 10.2989/18142320509504072

Smale, M.J. (2004)
Rajella barnardi IUCN 2012, IUCN Red List of Threatened Species, Version 2012.2, https://www.iucnredlist.org

Smale, M.J. (2004)
Anacanthobatis marmoratus IUCN 2012, IUCN Red List of Threatened Species, Version 2012.2, https://www.iucnredlist.org

Smale, M.J. (2004)
Anacanthobatis ori IUCN 2012, IUCN Red List of Threatened Species, Version 2012.2, https://www.iucnredlist.org

Smale, M.J. (2004)
Dipturus campbelli IUCN 2012, IUCN Red List of Threatened Species, Version 2012.2, https://www.iucnredlist.org

Smale, M.J. (2004)
Dipturus lanceorostratus IUCN 2012, IUCN Red List of Threatened Species, Version 2012.2, https://www.iucnredlist.org

Smale, M.J. (2004)
Dipturus pullopunctatus IUCN 2012, IUCN Red List of Threatened Species, Version 2012.2, https://www.iucnredlist.org

Smale, M.J. (2004)
Dipturus stenorhynchus IUCN 2012, IUCN Red List of Threatened Species, Version 2012.2, https://www.iucnredlist.org

Smale, M.J. (2004)
Neoraja stehmanni IUCN 2012, IUCN Red List of Threatened Species, Version 2012.2, https://www.iucnredlist.org

Smale, M.J. (2004)
Rajella caudaspinosa IUCN 2012, IUCN Red List of Threatened Species, Version 2012.2, https://www.iucnredlist.org

Smale, M.J. (2004)
Rajella dissimilis IUCN 2012, IUCN Red List of Threatened Species, Version 2012.2, https://www.iucnredlist.org

Smale, M.J. (2004)
Rajella leopardus IUCN 2012, IUCN Red List of Threatened Species, Version 2012.2, https://www.iucnredlist.org

Smale, M.J. (2004)
Rajella ravidula IUCN 2012, IUCN Red List of Threatened Species, Version 2012.2, https://www.iucnredlist.org

Smale, M.J. & Jones, R.T. & Correia, J.P. & Henningsen, A.D. & Crow, G.L. & Garner, R. (2004)
Chapter 39: Research on Elasmobranchs in Public Aquariums. In M. Smith, D. Warmolts, D. Thoney, & R. Hueter (Eds.), The Elasmobranch Husbandry Manual: Captive Care of Sharks, Rays and their Relatives (pp. 533–541). Special Publication of the Ohio Biological Survey

Henningsen, A.D. & Smale, M.J. & Garner, R. & Kinnunen, N. (2004)
Chapter 16: Reproduction, Embryonic Development, and Reproductive Physiology of Elasmobranchs. In M. Smith, D. Warmolts, D. Thoney, & R. Hueter (Eds.), The Elasmobranch Husbandry Manual: Captive Care of Sharks, Rays and their Relatives (pp. 227–236). Special Publication of the Ohio Biological Survey

Henningsen, A.D. & Smale, M.J. & Gordon, I. & Garner, R. & Marin-Osorno, R. & Kinnunen, N. (2004)
Chapter 17: Captive Breeding and Sexual Conflict in Elasmobranchs. In M. Smith, D. Warmolts, D. Thoney, & R. Hueter (Eds.), The Elasmobranch Husbandry Manual: Captive Care of Sharks, Rays and their Relatives (pp. 237–248). Special Publication of the Ohio Biological Survey

Smale, M.J. & Compagno, L.J.V. & Human, B.A. (2002)
First megamouth shark from the western Indian Ocean and South Africa. South African Journal of Science, 98(7/8), 349–350

Smale, M.J. (2002)
Occurrence of Carcharias taurus in nursery areas of the Eastern and Western Cape, South Africa Marine and Freshwater Research, 53(2), 551–556
DOI: 10.1071/MF01129

Voegeli, F.A. & Smale, M.J. & Webber, D.M. & Andrade, Y. & O'Dor, R.K. (2001)
Ultrasonic telemetry, tracking and automated monitoring technology for sharks. Environmental Biology of Fishes, 60(1–3), 267–282
DOI: 10.1023/A:1007682304720

Smale, M.J. & Sauer, W.H.H. & Roberts, M.J. (2001)
Behavioural interactions of predators and spawning chokka squid off South Africa: towards quantification. Marine Biology, 139(6), 1095–1105
DOI: 10.1007/s002270100664

Cliff, G. & Compagno, L.J.V. & Smale, M.J. & Van der Elst, R.P. & Wintner, S.P. (2000)
First records of white sharks, Carcharodon carcharias from Mauritius, Zanzibar, Madagascar and Kenya. South African Journal of Science, 96(7), 365–367

Watson, G. & Smale, M.J. (1999)
Age and growth of the shortnose spiny dogfish Squalus megalops from the Agulhas Bank, South Africa. South African Journal of Marine Science, 21(1), 9–18
DOI: 10.2989/025776199784126097

Smale, M.J. & Goosen, A.J.J. (1999)
Reproduction and feeding of spotted gully shark, Triakis megalopterus, off the Eastern Cape, South Africa. Fishery Bulletin, 97(4), 987–998

Watson, G. & Smale, M.J. (1998)
Reproductive biology of shortnose spiny dogfish, Squalus megalops, from the Agulhas Bank, South Africa. Marine and Freshwater Research, 49(7), 695–703
DOI: 10.1071/MF97255

Smale, M.J. & Cliff, G. (1998)
Cephalopods in the diets of four shark species (Galeocerdo cuvier, Sphyrna lewini, S. zygaena and S. mokarran) from KwaZulu-Natal, South Africa. South African Journal of Marine Science, 20(1), 241–253
DOI: 10.2989/025776198784126610

Beckley, L.E. & Cliff, G.M. & Smale, M.J. & Compagno, L.J.V. (1997)
Recent strandings and sightings of whale sharks in South Africa. Environmental Biology of Fishes, 50(3), 343–348
DOI: 10.1023/A:1007355709632

Smale, M.J. & Heemstra, P.C. (1997)
First record of albinism in the great white shark, Carcharodon carcharias (Linnaeus, 1758). South African Journal of Science, 93(5), 243–245

Smale, M.J. & Compagno, L.J.V. (1997)
Life history and diet of two southern African smoothhound sharks, Mustelus mustelus (Linnaeus, 1758) and Mustelus palumbes Smith, 1957 (Pisces: Triakidae). South African Journal of Marine Science, 18(1), 229–248
DOI: 10.2989/025776197784160992

Goosen, A.J.J. & Smale, M.J. (1997)
A preliminary study of the age and Size of the smoothhound shark Mustelus mustelus (Triakidae). South African Journal of Marine Science, 18(1), 85–91
DOI: 10.2989/025776197784161072

Smale, M.J. & Sauer, W.H.H. & Hanlon, R.T. (1995)
Attempted ambush predation on spawning squids Loligo vulgaris reynaudii by benthic pyjama sharks, Poroderma africanum, off South Africa. Journal of the Marine Biological Association of the United Kingdom, 75(3), 739–742
DOI: 10.1017/S002531540003914X

Oldewage, W.H. & Smale, M.J. (1993)
Occurrence of piscine parasitic copepods (Crustacea) on sharks taken mainly off Cape Recife, South Africa. South African Journal of Marine Science, 13(1), 309–312
DOI: 10.2989/025776193784287310

Smale, M.J. & Cowley, P.D. (1992)
The feeding ecology of skates (Batoidea: Rajidae) off the cape south coast, South Africa. South African Journal of Marine Science, 12(1), 823–834
DOI: 10.2989/02577619209504744

Smale, M.J. (1991)
Occurrence and feeding of three shark species, Carcharhinus brachyurus, C. obscurus and Sphyrna zygaena, on the eastern Cape Coast of South Africa. South African Journal of Marine Science, 11(1), 31–42
DOI: 10.2989/025776191784287808

Compagno, L.J.V. & Ebert, D.A. & Smale, M.J. (1989)
Guide to the sharks and rays of Southern Africa. New Holland Ltd., London 1989

Compagno, L.J.V. & Smale, M.J. (1986)
Recent records of four warm-water elasmobranchs from the eastern Cape Province, South Africa. South African Journal of Marine Science, 4(1), 11–15, fig
DOI: 10.2989/025776186784461873

Compagno, L.J.V. & Smale, M.J. (1985)
Paragaleus leucolomatus, a new shark from South Africa, with notes on the systematics of hemigaleid sharks (Carcharhiniformes: Hemigaleidae). J.L.B. Smith Institute of Ichthyology Special Publication, 37: 1–21