Arlyza, I.S. & Shen, K.-N. & Solihin, D.D. & Soedharma, D. & Berrebi, P. & Borsa, P. (2013)
Species boundaries in the Himantura uarnak species complex (Myliobatiformes: Dasyatidae).
Molecular Phylogenetics and Evolution, 66(1), 429–435 DOI: 10.1016/j.ympev.2012.09.023