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Haemogregarina delagei (2)
Laveran & Mesnil, 1901
  • Amblyraja radiata (Donovan, 1808) (Rajiformes: Rajidae)
    • Khan, R.A. et.al. (1991) [21433] (used host: Raja radiata )
  • Bathyraja spinicauda (Jensen, 1914) (Rajiformes: Arhynchobatidae)
    • Khan, R.A. et.al. (1991) [21433] (used host: Raja spinicauda )
  • Raja microocellata Montagu, 1818 (Rajiformes: Rajidae)
    • Aragort, W. et.al. (2005) [21437]
Original description species: Laveran & Mesnil, 1901
Genus: Haemogregarina Danilewsky, 1885
Family: Haemogregarinidae Léger, 1911
Suborder: Adeleorina
Order: Eucoccidiorida
Subclass: Coccidiasina
Haemohormidium sp. (1)

  • Centroscyllium fabricii (Reinhardt, 1825) (Squaliformes: Etmopteridae)
Genus: Haemohormidium Henry, 1910
Family: Haemohormidiidae Levine, 1984
Order: Piroplasmorida
Halysioncum arafurense (1)
Ivanov & Caira, 2013
  • Aetomylaeus nichofii (Bloch & Schneider, 1801) (Myliobatiformes: Myliobatidae)
    • Ivanov, V.A. et.al. (2013) [19639] (used host: Aetomylaeus cf. nichofii )
Original description species: Ivanov & Caira, 2013
Genus: Halysioncum Caira, Marques, Jensen, Kuchta & Ivanov, 2013
Family: Echinobothriidae Perrier, 1897
Order: Diphyllidea
Subclass: Eucestoda
Halysioncum boisii (3)
(Southwell, 1911)
  • Aetobatus narinari (Euphrasen, 1790) (Myliobatiformes: Aetobatidae)
    • Southwell, T. (1911) [17129] ( used parasite: Echinobothrium boisii)
    • Tyler, G.A. (2006) [16430] ( used parasite: Echinobothrium boisii)
    • Caira, J.N. et.al. (2013) [19536]
Synonyms: Echinobothrium boisii
Original description species: (Southwell, 1911)
Genus: Halysioncum Caira, Marques, Jensen, Kuchta & Ivanov, 2013
Family: Echinobothriidae Perrier, 1897
Order: Diphyllidea
Subclass: Eucestoda
Halysioncum bonasum (3)
(Williams & Campbell, 1980)
  • Rhinoptera bonasus (Mitchill, 1815) (Myliobatiformes: Rhinopteridae)
    • Williams, A.D. et.al. (1980) [7577] ( used parasite: Echinobothrium bonasum)
    • Tyler, G.A. (2006) [16430] ( used parasite: Echinobothrium bonasum)
    • Caira, J.N. et.al. (2013) [19536]
  • Rhinoptera steindachneri Evermann & Jenkins, 1891 (Myliobatiformes: Rhinopteridae)
    • Caira, J.N. et.al. (2013) [19536] (used host: Rhinoptera cf. steindachneri )
Synonyms: Echinobothrium bonasum
Original description species: (Williams & Campbell, 1980)
Genus: Halysioncum Caira, Marques, Jensen, Kuchta & Ivanov, 2013
Family: Echinobothriidae Perrier, 1897
Order: Diphyllidea
Subclass: Eucestoda
Halysioncum californiense (3)
(Ivanov & Campbell, 1998)
  • Platyrhinoidis triseriata (Jordan & Gilbert, 1880) (Rhinopristiformes: Platyrhinidae)
    • Ivanov, V.A. et.al. (1998) [16327] ( used parasite: Echinobothrium californiense)
    • Tyler, G.A. (2006) [16430] ( used parasite: Echinobothrium californiense)
    • Caira, J.N. et.al. (2013) [19536]
Synonyms: Echinobothrium californiense
Original description species: (Ivanov & Campbell, 1998)
Genus: Halysioncum Caira, Marques, Jensen, Kuchta & Ivanov, 2013
Family: Echinobothriidae Perrier, 1897
Order: Diphyllidea
Subclass: Eucestoda
Halysioncum euzeti (4)
(Campbell & Carvajal, 1980)
  • Sympterygia lima (Poeppig, 1835) (Rajiformes: Arhynchobatidae)
    • Campbell, R.A. et.al. (1980) [16285] (used host: Psammobatis lima | used parasite: Echinobothrium euzeti)
    • Tyler, G.A. (2006) [16430] ( used parasite: Echinobothrium euzeti)
    • Caira, J.N. et.al. (2013) [19536]
    • Alves, P.V. et.al. (2017) [25154]
Synonyms: Echinobothrium euzeti
Original description species: (Campbell & Carvajal, 1980)
Genus: Halysioncum Caira, Marques, Jensen, Kuchta & Ivanov, 2013
Family: Echinobothriidae Perrier, 1897
Order: Diphyllidea
Subclass: Eucestoda
Halysioncum fautleyae (4)
(Tyler & Caira, 1999)
  • Myliobatis californica Gill, 1865 (Myliobatiformes: Myliobatidae)
    • Tyler, G.A. et.al. (1999) [7627] ( used parasite: Echinobothrium fautleyae)
    • Caira, J.N. et.al. (2013) [19536]
    • IVan Merlo-Serna, A. et.al. (2016) [23771] ( used parasite: Echinobothrium fautleyae)
  • Myliobatis longirostris Applegate & Fitch, 1964 (Myliobatiformes: Myliobatidae)
    • Caira, J.N. et.al. (2013) [19536]
  • Rhinoptera steindachneri Evermann & Jenkins, 1891 (Myliobatiformes: Rhinopteridae)
    • Tyler, G.A. et.al. (1999) [7627] ( used parasite: Echinobothrium fautleyae)
    • Tyler, G.A. (2006) [16430] ( used parasite: Echinobothrium fautleyae)
    • IVan Merlo-Serna, A. et.al. (2016) [23771] ( used parasite: Echinobothrium fautleyae)
Synonyms: Echinobothrium fautleyae
Original description species: (Tyler & Caira, 1999)
Genus: Halysioncum Caira, Marques, Jensen, Kuchta & Ivanov, 2013
Family: Echinobothriidae Perrier, 1897
Order: Diphyllidea
Subclass: Eucestoda
Halysioncum gibsoni (1)
Ivanov & Caira, 2013
  • Aetomylaeus maculatus (Gray, 1834) (Myliobatiformes: Myliobatidae)
    • Ivanov, V.A. et.al. (2013) [19639]
Original description species: Ivanov & Caira, 2013
Genus: Halysioncum Caira, Marques, Jensen, Kuchta & Ivanov, 2013
Family: Echinobothriidae Perrier, 1897
Order: Diphyllidea
Subclass: Eucestoda
Halysioncum hoffmanorum (4)
(Tyler, 2001)
  • Urobatis concentricus Osburn & Nichols, 1916 (Myliobatiformes: Urotrygonidae)
    • Tyler, G.A. (2001) [16203] ( used parasite: Echinobothrium hoffmanorum)
    • Tyler, G.A. (2006) [16430] ( used parasite: Echinobothrium hoffmanorum)
    • Caira, J.N. et.al. (2013) [19536]
    • IVan Merlo-Serna, A. et.al. (2016) [23771] ( used parasite: Echinobothrium hoffmanorum)
  • Urobatis halleri (Cooper, 1863) (Myliobatiformes: Urotrygonidae)
    • Tyler, G.A. (2001) [16203] ( used parasite: Echinobothrium hoffmanorum)
    • Tyler, G.A. (2006) [16430] ( used parasite: Echinobothrium hoffmanorum)
    • Caira, J.N. et.al. (2013) [19536]
    • IVan Merlo-Serna, A. et.al. (2016) [23771] (used host: Urolophus halleri | used parasite: Echinobothrium hoffmanorum)
  • Urobatis maculatus Garman, 1913 (Myliobatiformes: Urotrygonidae)
    • Tyler, G.A. (2001) [16203] ( used parasite: Echinobothrium hoffmanorum)
    • Tyler, G.A. (2006) [16430] ( used parasite: Echinobothrium hoffmanorum)
    • Caira, J.N. et.al. (2013) [19536]
    • IVan Merlo-Serna, A. et.al. (2016) [23771] (used host: Urolophus maculatus | used parasite: Echinobothrium hoffmanorum)
Synonyms: Echinobothrium hoffmanorum
Original description species: (Tyler, 2001)
Genus: Halysioncum Caira, Marques, Jensen, Kuchta & Ivanov, 2013
Family: Echinobothriidae Perrier, 1897
Order: Diphyllidea
Subclass: Eucestoda
Halysioncum kishiense (1)
Moghadam & Meraji Masouleh Haseli, 2014
  • Aetomylaeus nichofii (Bloch & Schneider, 1801) (Myliobatiformes: Myliobatidae)
    • Moghadam, M. et.al. (2014) [21001] (used host: Aetomylaeus cf. nichofii )
Original description species: Moghadam & Meraji Masouleh Haseli, 2014
Genus: Halysioncum Caira, Marques, Jensen, Kuchta & Ivanov, 2013
Family: Echinobothriidae Perrier, 1897
Order: Diphyllidea
Subclass: Eucestoda
Halysioncum megacanthum (5)
(Ivanov & Campbell, 1998)
  • Myliobatis goodei Garman, 1885 (Myliobatiformes: Myliobatidae)
    • Ivanov, V.A. et.al. (1998) [15941] ( used parasite: Echinobothrium megacanthum)
    • Tyler, G.A. (2006) [16430] ( used parasite: Echinobothrium megacanthum)
    • Caira, J.N. et.al. (2013) [19536]
    • Alves, P.V. et.al. (2017) [25154]
    • Franzese, S. et.al. (2023) [31906]
Synonyms: Echinobothrium megacanthum
Original description species: (Ivanov & Campbell, 1998)
Genus: Halysioncum Caira, Marques, Jensen, Kuchta & Ivanov, 2013
Family: Echinobothriidae Perrier, 1897
Order: Diphyllidea
Subclass: Eucestoda
Halysioncum mexicanum (4)
(Tyler & Caira, 1999)
  • Myliobatis californica Gill, 1865 (Myliobatiformes: Myliobatidae)
    • Tyler, G.A. et.al. (1999) [7627] ( used parasite: Echinobothrium mexicanum)
    • Tyler, G.A. (2006) [16430] ( used parasite: Echinobothrium mexicanum)
    • Caira, J.N. et.al. (2013) [19536]
    • IVan Merlo-Serna, A. et.al. (2016) [23771] ( used parasite: Echinobothrium mexicanum)
  • Myliobatis longirostris Applegate & Fitch, 1964 (Myliobatiformes: Myliobatidae)
    • Tyler, G.A. et.al. (1999) [7627] ( used parasite: Echinobothrium mexicanum)
    • Tyler, G.A. (2006) [16430] ( used parasite: Echinobothrium mexicanum)
    • IVan Merlo-Serna, A. et.al. (2016) [23771] ( used parasite: Echinobothrium mexicanum)
  • Rhinoptera steindachneri Evermann & Jenkins, 1891 (Myliobatiformes: Rhinopteridae)
    • Caira, J.N. et.al. (2013) [19536]
Synonyms: Echinobothrium mexicanum
Original description species: (Tyler & Caira, 1999)
Genus: Halysioncum Caira, Marques, Jensen, Kuchta & Ivanov, 2013
Family: Echinobothriidae Perrier, 1897
Order: Diphyllidea
Subclass: Eucestoda
Halysioncum nataliae (2)
(Kuchta & Caira, 2010)
  • Pastinachus solocirostris Last, Manjaji & Yearsley, 2005 (Myliobatiformes: Dasyatidae)
    • Kuchta, R. et.al. (2010) [12201] ( used parasite: Echinobothrium nataliae)
    • Caira, J.N. et.al. (2013) [19536]
Synonyms: Echinobothrium nataliae
Original description species: (Kuchta & Caira, 2010)
Genus: Halysioncum Caira, Marques, Jensen, Kuchta & Ivanov, 2013
Family: Echinobothriidae Perrier, 1897
Order: Diphyllidea
Subclass: Eucestoda
Halysioncum pigmentatum (5)
(Ostrowski De Núnez, 1971)
  • Zapteryx brevirostris (Müller & Henle, 1841) (Rhinopristiformes: Trygonorrhinidae)
    • Ostrowski de Nuñez, M. (1971) [16365] ( used parasite: Echinobothrium pigmentatum)
    • Tyler, G.A. (2006) [16430] ( used parasite: Echinobothrium pigmentatum)
    • Caira, J.N. et.al. (2013) [19536]
    • Alves, P.V. et.al. (2017) [25154]
    • Franzese, S. et.al. (2023) [31906]
Synonyms: Echinobothrium pigmentatum
Original description species: (Ostrowski De Núnez, 1971)
Genus: Halysioncum Caira, Marques, Jensen, Kuchta & Ivanov, 2013
Family: Echinobothriidae Perrier, 1897
Order: Diphyllidea
Subclass: Eucestoda
Halysioncum raschii (3)
(Campbell & Andrade, 1997)
  • Bathyraja taranetzi (Dolganov, 1983) (Rajiformes: Arhynchobatidae)
    • Campbell, R.A. et.al. (1997) [7579] (used host: Rhinoraja longi | used parasite: Echinobothrium raschii)
    • Tyler, G.A. (2006) [16430] (used host: Rhinoraja longi | used parasite: Echinobothrium raschii)
    • Caira, J.N. et.al. (2013) [19536] (used host: Rhinoraja longi )
Synonyms: Echinobothrium raschii
Original description species: (Campbell & Andrade, 1997)
Genus: Halysioncum Caira, Marques, Jensen, Kuchta & Ivanov, 2013
Family: Echinobothriidae Perrier, 1897
Order: Diphyllidea
Subclass: Eucestoda
Halysioncum rayallemangi (4)
(Tyler, 2001)
  • Pseudobatos leucorhynchus (Günther, 1866) (Rhinopristiformes: Rhinobatidae)
    • Tyler, G.A. (2001) [16203] (used host: Rhinobatos leucorhynchus | used parasite: Echinobothrium rayallemangi)
    • Tyler, G.A. (2006) [16430] (used host: Rhinobatos leucorhynchus | used parasite: Echinobothrium rayallemangi)
    • Caira, J.N. et.al. (2013) [19536] (used host: Rhinobatos leucorhynchus )
    • IVan Merlo-Serna, A. et.al. (2016) [23771] (used host: Rhinobatos leucorhynchus | used parasite: Echinobothrium rayallemangi)
Synonyms: Echinobothrium rayallemangi
Original description species: (Tyler, 2001)
Genus: Halysioncum Caira, Marques, Jensen, Kuchta & Ivanov, 2013
Family: Echinobothriidae Perrier, 1897
Order: Diphyllidea
Subclass: Eucestoda
Halysioncum reginae (2)
(Kuchta & Caira, 2010)
  • Pastinachus ater (Macleay, 1883) (Myliobatiformes: Dasyatidae)
    • Caira, J.N. et.al. (2013) [19536] (used host: Pastinachus cf. atrus )
  • Pastinachus sephen (Fabricius in Niebuhr, 1775) (Myliobatiformes: Dasyatidae)
    • Kuchta, R. et.al. (2010) [12201] (used host: Pastinachus cf. sephen | used parasite: Echinobothrium reginae)
Synonyms: Echinobothrium reginae
Original description species: (Kuchta & Caira, 2010)
Genus: Halysioncum Caira, Marques, Jensen, Kuchta & Ivanov, 2013
Family: Echinobothriidae Perrier, 1897
Order: Diphyllidea
Subclass: Eucestoda
Halysioncum rhinoptera (2)
(Shipley & Hornell, 1906)
  • Rhinoptera javanica Müller & Henle, 1841 (Myliobatiformes: Rhinopteridae)
    • Tyler, G.A. (2006) [16430] ( used parasite: Echinobothrium rhinoptera)
    • Caira, J.N. et.al. (2013) [19536]
Synonyms: Echinobothrium rhinoptera
Original description species: (Shipley & Hornell, 1906)
Genus: Halysioncum Caira, Marques, Jensen, Kuchta & Ivanov, 2013
Family: Echinobothriidae Perrier, 1897
Order: Diphyllidea
Subclass: Eucestoda
Halysioncum sp. (1)

  • Myliobatis aquila (Linnaeus, 1758) (Myliobatiformes: Myliobatidae)
    • Caira, J.N. et.al. (2013) [19536]
  • Pastinachus ater (Macleay, 1883) (Myliobatiformes: Dasyatidae)
    • Caira, J.N. et.al. (2013) [19536] (used host: Pastinachus atrus )
  • Pteromylaeus bovinus (Geoffroy Saint-Hilaire, 1817) (Myliobatiformes: Myliobatidae)
    • Caira, J.N. et.al. (2013) [19536]
  • Rhinoptera bonasus (Mitchill, 1815) (Myliobatiformes: Rhinopteridae)
    • Caira, J.N. et.al. (2013) [19536] (used host: Rhinoptera cf. bonasus )
  • Rhinoptera neglecta Ogilby, 1912 (Myliobatiformes: Rhinopteridae)
    • Caira, J.N. et.al. (2013) [19536]
Genus: Halysioncum Caira, Marques, Jensen, Kuchta & Ivanov, 2013
Family: Echinobothriidae Perrier, 1897
Order: Diphyllidea
Subclass: Eucestoda
Halysioncum vojtai (2)
(Kuchta & Caira, 2010)
  • Pastinachus gracilicaudus Last & Manjaji-Matsumoto, 2010 (Myliobatiformes: Dasyatidae)
    • Caira, J.N. et.al. (2013) [19536]
Synonyms: Echinobothrium vojtai
Original description species: (Kuchta & Caira, 2010)
Genus: Halysioncum Caira, Marques, Jensen, Kuchta & Ivanov, 2013
Family: Echinobothriidae Perrier, 1897
Order: Diphyllidea
Subclass: Eucestoda
Halysiorhynchus macrocephalus (7)
(Shipley & Hornell, 1906)
  • Brevitrygon walga (Müller & Henle, 1841) (Myliobatiformes: Dasyatidae)
    • Palm, H.W. (2004) [16112] (used host: Himantura walga )
  • Gymnura micrura (Bloch & Schneider, 1801) (Myliobatiformes: Gymnuridae)
  • Himantura uarnak (Forsskål, 1775) (Myliobatiformes: Dasyatidae)
  • Maculabatis gerrardi (Gray, 1851) (Myliobatiformes: Dasyatidae)
    • Schaeffner, B.C. et.al. (2014) [21816] (used host: Himantura gerrardi )
  • Neotrygon kuhlii (Müller & Henle, 1841) (Myliobatiformes: Dasyatidae)
    • Palm, H.W. (2004) [16112] (used host: Dasyatis kuhlii )
  • Pastinachus sephen (Fabricius in Niebuhr, 1775) (Myliobatiformes: Dasyatidae)
    • Palm, H.W. (2004) [16112]
    • Palm, H.W. et.al. (2009) [15747]
    • Haseli, M. et.al. (2010) [15730] (used host: Pastinachus cf. sephen )
  • Pateobatis uarnacoides (Bleeker, 1852) (Myliobatiformes: Dasyatidae)
    • Schaeffner, B.C. et.al. (2014) [21816] (used host: Himantura uarnacoides )
  • Rhina ancylostoma Bloch & Schneider, 1801 (Rhinopristiformes: Rhinidae)
    • Schaeffner, B.C. et.al. (2014) [21816]
    • Shafiei, S. et.al. (2019) [27290]
  • Rhynchobatus djiddensis (Forsskål, 1775) (Rhinopristiformes: Rhinidae)
  • Rhynchobatus laevis (Bloch & Schneider, 1801) (Rhinopristiformes: Rhinidae)
    • Schaeffner, B.C. et.al. (2014) [21816] (used host: Rhynchobatus cf. laevis )
Synonyms: Tetrarhynchus macrocephalus
Original description species: (Shipley & Hornell, 1906)
Genus: Halysiorhynchus Pintner, 1913
Family: Mixodigmatidae Dailey & Vogelbein, 1982
Superfamily: Eutetrarhynchoidea
Suborder: Trypanobatoida
Order: Trypanorhyncha
Subclass: Eucestoda
Healyum harenamica (3)
Jensen, 2001
  • Mobula mobular (Bonnaterre, 1788) (Myliobatiformes: Mobulidae)
    • Jensen, K. (2001) [7588] (used host: Mobula japanica )
    • Jensen, K. (2005) [16338] (used host: Mobula japanica )
    • IVan Merlo-Serna, A. et.al. (2016) [23771] (used host: Mobula japanica )
Original description species: Jensen, 2001
Genus: Healyum Jensen, 2001
Family: Lecanicephalidea incertae sedis Braun, 1900
Order: Lecanicephalidea
Subclass: Eucestoda
Healyum pulvis (3)
Jensen, 2001
  • Mobula mobular (Bonnaterre, 1788) (Myliobatiformes: Mobulidae)
    • Jensen, K. (2001) [7588] (used host: Mobula japanica )
    • Jensen, K. (2005) [16338] (used host: Mobula japanica )
    • IVan Merlo-Serna, A. et.al. (2016) [23771] (used host: Mobula japanica )
Original description species: Jensen, 2001
Genus: Healyum Jensen, 2001
Family: Lecanicephalidea incertae sedis Braun, 1900
Order: Lecanicephalidea
Subclass: Eucestoda
Helicometrina nimia (1)
Linton, 1910
  • Prionace glauca (Linnaeus, 1758) (Carcharhiniformes: Carcharhinidae)
    • IVan Merlo-Serna, A. et.al. (2016) [23771]
Original description species: Linton, 1910
Genus: Helicometrina Linton, 1910
Family: Opecoelidae Ozaki, 1925
Superfamily: Allocreadioidea
Suborder: Xiphidiata
Order: Plagiorchiida
Subclass: Digenea
Heliconema psammobatidus (1)
Threlfall & Carvajal, 1984
  • Sympterygia lima (Poeppig, 1835) (Rajiformes: Arhynchobatidae)
    • Pereira, F.B. et.al. (2023) [31368]
Original description species: Threlfall & Carvajal, 1984
Genus: Heliconema Travassos, 1919
Family: Physalopteridae Railliet, 1893
Superfamily: Physalopteroidea
Suborder: Spirurina
Order: Spirurida
Heliconema sp. (1)

  • Cephalurus cephalus (Gilbert, 1892) (Carcharhiniformes: Pentanchidae)
    • Morales-Ávila, J.R. et.al. (2023) [32170]
Genus: Heliconema Travassos, 1919
Family: Physalopteridae Railliet, 1893
Superfamily: Physalopteroidea
Suborder: Spirurina
Order: Spirurida
Heliocotyle ewingi (1)
Chisholm & Whittington, 2000
  • Myliobatis tenuicaudatus Hector, 1877 (Myliobatiformes: Myliobatidae)
    • Chisholm, L.A. et.al. (2000) [9434] (used host: Myliobatis australis )
Original description species: Chisholm & Whittington, 2000
Genus: Heliocotyle Neifar, Euzet & Ben Hassine, 1999
Family: Monocotylidae Taschenberg, 1879
Order: Monocotylidea
Subclass: Monopisthocotylea
Heliocotyle kartasi (3)
Neifar & Euzet & Ben Hassine, 1999
  • Pteromylaeus bovinus (Geoffroy Saint-Hilaire, 1817) (Myliobatiformes: Myliobatidae)
    • Neifar, L. et.al. (1999) [21211]
    • Derbel, H. et.al. (2022) [31118] (used host: Aetomylaeus bovinus )
    • Vaughan, D.B. et.al. (2024) [33260] (used host: Aetomylaeus bovinus )
Original description species: Neifar & Euzet & Ben Hassine, 1999
Genus: Heliocotyle Neifar, Euzet & Ben Hassine, 1999
Family: Monocotylidae Taschenberg, 1879
Order: Monocotylidea
Subclass: Monopisthocotylea
Hemionchos maior (2)
Campbell & Beveridge, 2006
  • Mobula mobular (Bonnaterre, 1788) (Myliobatiformes: Mobulidae)
    • Campbell, R.A. et.al. (2006) [9451] (used host: Mobula japanica )
    • IVan Merlo-Serna, A. et.al. (2016) [23771] (used host: Mobula japanica )
Original description species: Campbell & Beveridge, 2006
Genus: Hemionchos Campbell & Beveridge, 2006
Family: Eutetrarhynchidae Guiart, 1927
Superfamily: Eutetrarhynchoidea
Suborder: Trypanobatoida
Order: Trypanorhyncha
Subclass: Eucestoda
Hemionchos mobulae (2)
Campbell & Beveridge, 2006
  • Mobula mobular (Bonnaterre, 1788) (Myliobatiformes: Mobulidae)
    • Campbell, R.A. et.al. (2006) [9451] (used host: Mobula japanica )
    • IVan Merlo-Serna, A. et.al. (2016) [23771] (used host: Mobula japanica )
  • Mobula munkiana Notarbartolo Di Sciara, 1987 (Myliobatiformes: Mobulidae)
    • Campbell, R.A. et.al. (2006) [9451]
    • IVan Merlo-Serna, A. et.al. (2016) [23771]
Original description species: Campbell & Beveridge, 2006
Genus: Hemionchos Campbell & Beveridge, 2006
Family: Eutetrarhynchidae Guiart, 1927
Superfamily: Eutetrarhynchoidea
Suborder: Trypanobatoida
Order: Trypanorhyncha
Subclass: Eucestoda
Hemionchos striatus (2)
Campbell & Beveridge, 2006
  • Mobula mobular (Bonnaterre, 1788) (Myliobatiformes: Mobulidae)
    • Campbell, R.A. et.al. (2006) [9451] (used host: Mobula japanica )
    • IVan Merlo-Serna, A. et.al. (2016) [23771] (used host: Mobula japanica )
  • Mobula thurstoni (Lloyd, 1908) (Myliobatiformes: Mobulidae)
    • Campbell, R.A. et.al. (2006) [9451]
    • IVan Merlo-Serna, A. et.al. (2016) [23771]
  • Myliobatis californica Gill, 1865 (Myliobatiformes: Myliobatidae)
    • IVan Merlo-Serna, A. et.al. (2016) [23771]
Original description species: Campbell & Beveridge, 2006
Genus: Hemionchos Campbell & Beveridge, 2006
Family: Eutetrarhynchidae Guiart, 1927
Superfamily: Eutetrarhynchoidea
Suborder: Trypanobatoida
Order: Trypanorhyncha
Subclass: Eucestoda
Hemipristicola gunterae (2)
Cutmore, Theiss, Bennett & Cribb, 2011
  • Hemipristis elongata (Klunzinger, 1871) (Carcharhiniformes: Hemigaleidae)
    • Cutmore, S.C. et.al. (2011) [14277]
    • Cutmore, S.C. et.al. (2017) [25840]
Original description species: Cutmore, Theiss, Bennett & Cribb, 2011
Genus: Hemipristicola Cutmore, Theiss, Bennett & Cribb, 2011
Family: Phyllobothriidae Braun, 1900
Order: Tetraphyllidea
Subclass: Eucestoda
Hemiurus levinseni (1)
Odhner, 1905
  • Dipturus laevis (Mitchill, 1818) (Rajiformes: Rajidae)
    • Myers, B.J. (1959) [12575] (used host: Raja laevis )
  • Squalus acanthias Linnaeus, 1758 (Squaliformes: Squalidae)
Original description species: Odhner, 1905
Genus: Hemiurus Rudolphi, 1809
Family: Hemiuridae Looss, 1899
Superfamily: Hemiuroidea
Suborder: Hemiurata
Order: Plagiorchiida
Subclass: Digenea
Hepatoxylon megacephalum (3)
(Rudolphi, 1819)
  • Carcharhinus obscurus (Lesueur, 1818) (Carcharhiniformes: Carcharhinidae)
  • Carcharodon carcharias (Linnaeus, 1758) (Lamniformes: Lamnidae)
  • Dalatias licha (Bonnaterre, 1788) (Squaliformes: Dalatiidae)
  • Deania calceus (Lowe, 1839) (Squaliformes: Centrophoridae)
    • Waterman, P.B. et.al. (1991) [21481] (used host: Deania calcea )
    • Palm, H.W. (2004) [16112] (used host: Deania calcea )
  • Dipturus oxyrinchus (Linnaeus, 1758) (Rajiformes: Rajidae)
  • Etmopterus spinax (Linnaeus, 1758) (Squaliformes: Etmopteridae)
  • Galeorhinus galeus (Linnaeus, 1758) (Carcharhiniformes: Triakidae)
    • Hewitt, G.C. et.al. (1972) [16602] (used host: Galeorhinus australis )
    • Waterman, P.B. et.al. (1991) [21481]
    • Palm, H.W. (2004) [16112]
  • Galeus melastomus Rafinesque, 1810 (Carcharhiniformes: Pentanchidae)
  • Heptranchias perlo (Bonnaterre, 1788) (Hexanchiformes: Hexanchidae)
  • Hexanchus griseus (Bonnaterre, 1788) (Hexanchiformes: Hexanchidae)
  • Isurus oxyrinchus Rafinesque, 1810 (Lamniformes: Lamnidae)
  • Mustelus mustelus (Linnaeus, 1758) (Carcharhiniformes: Triakidae)
  • Notorynchus cepedianus (Péron, 1807) (Hexanchiformes: Hexanchidae)
  • Prionace glauca (Linnaeus, 1758) (Carcharhiniformes: Carcharhinidae)
  • Raja clavata Linnaeus, 1758 (Rajiformes: Rajidae)
  • Scyliorhinus canicula (Linnaeus, 1758) (Carcharhiniformes: Scyliorhinidae)
  • Scyliorhinus stellaris (Linnaeus, 1758) (Carcharhiniformes: Scyliorhinidae)
  • Sphyrna zygaena (Linnaeus, 1758) (Carcharhiniformes: Sphyrnidae)
  • Squalus acanthias Linnaeus, 1758 (Squaliformes: Squalidae)
  • Squalus megalops (Macleay, 1881) (Squaliformes: Squalidae)
  • Squatina australis Regan, 1906 (Squatiniformes: Squatinidae)
  • Torpedo marmorata Risso, 1810 (Torpediniformes: Torpedinidae)
Original description species: (Rudolphi, 1819)
Genus: Hepatoxylon Bosc, 1811
Family: Sphyriocephalidae Pinter, 1913
Superfamily: Tentacularioidea
Suborder: Trypanobatoida
Order: Trypanorhyncha
Subclass: Eucestoda
Hepatoxylon sp. (3)

  • Apristurus nasutus De Buen, 1959 (Carcharhiniformes: Pentanchidae)
    • Espínola-Novelo, J.F. et.al. (2018) [26778]
  • Etmopterus granulosus (Günther, 1880) (Squaliformes: Etmopteridae)
    • Espínola-Novelo, J.F. et.al. (2018) [26778]
  • Prionace glauca (Linnaeus, 1758) (Carcharhiniformes: Carcharhinidae)
  • Sphyrna tiburo (Linnaeus, 1758) (Carcharhiniformes: Sphyrnidae)
Genus: Hepatoxylon Bosc, 1811
Family: Sphyriocephalidae Pinter, 1913
Superfamily: Tentacularioidea
Suborder: Trypanobatoida
Order: Trypanorhyncha
Subclass: Eucestoda
Hepatoxylon trichiuri (13)
(Holten, 1802)
  • Alopias superciliosus (Lowe, 1841) (Lamniformes: Alopiidae)
  • Alopias vulpinus (Bonnaterre, 1788) (Lamniformes: Alopiidae)
  • Carcharhinus plumbeus (Nardo, 1827) (Carcharhiniformes: Carcharhinidae)
  • Carcharodon carcharias (Linnaeus, 1758) (Lamniformes: Lamnidae)
  • Centroscymnus coelolepis Barbosa Du Bocage & De Brito Capello, 1864 (Squaliformes: Somniosidae)
  • Centroscymnus owstonii Garman, 1906 (Squaliformes: Somniosidae)
  • Chlamydoselachus anguineus Garman, 1884 (Hexanchiformes: Chlamydoselachidae)
  • Dalatias licha (Bonnaterre, 1788) (Squaliformes: Dalatiidae)
  • Deania calceus (Lowe, 1839) (Squaliformes: Centrophoridae)
    • Hewitt, G.C. et.al. (1972) [16602] (used host: Deania calcea )
    • Waterman, P.B. et.al. (1991) [21481] (used host: Deania calcea )
    • Palm, H.W. (2004) [16112] (used host: Deania calcea )
  • Galeorhinus galeus (Linnaeus, 1758) (Carcharhiniformes: Triakidae)
  • Galeus melastomus Rafinesque, 1810 (Carcharhiniformes: Pentanchidae)
  • Hexanchus griseus (Bonnaterre, 1788) (Hexanchiformes: Hexanchidae)
  • Isurus oxyrinchus Rafinesque, 1810 (Lamniformes: Lamnidae)
  • Lamna nasus (Bonnaterre, 1788) (Lamniformes: Lamnidae)
  • Mustelus mustelus (Linnaeus, 1758) (Carcharhiniformes: Triakidae)
  • Prionace glauca (Linnaeus, 1758) (Carcharhiniformes: Carcharhinidae)
    • Yamaguti, S. (1934) [16458] ( used parasite: Dibothriorhynchus squali)
    • Yamaguti, S. (1934) [16458] (used host: Isurus glaucus | used parasite: Dibothriorhynchus squali)
    • Pappas, P.W. (1970) [23382] ( used parasite: Hepatoxylon squali)
    • Hewitt, G.C. et.al. (1972) [16602]
    • Carvajal, J. (1974) [16182]
    • Waterman, P.B. et.al. (1991) [21481]
    • Campbell, R.A. et.al. (1998) [9432]
    • Henderson, A.C. et.al. (2002) [9354]
    • Cousin, J.C.B. et.al. (2003) [21423]
    • Knoff, M. et.al. (2004) [19882]
    • Palm, H.W. (2004) [16112]
    • Beveridge, I. et.al. (2014) [21718]
    • Alves, P.V. et.al. (2017) [25154]
  • Scyliorhinus canicula (Linnaeus, 1758) (Carcharhiniformes: Scyliorhinidae)
  • Somniosus microcephalus (Bloch & Schneider, 1801) (Squaliformes: Somniosidae)
  • Somniosus pacificus Bigelow & Schroeder, 1944 (Squaliformes: Somniosidae)
  • Squalus acanthias Linnaeus, 1758 (Squaliformes: Squalidae)
  • Tetronarce nobiliana (Bonaparte, 1835) (Torpediniformes: Torpedinidae)
    • Hewitt, G.C. et.al. (1972) [16602] (used host: Torpedo fairchildi )
    • Palm, H.W. (2004) [16112] (used host: Torpedo fairchildi )
Synonyms: Dibothriorhynchus squali, Hepatoxylon squali, Hirudo squali
Original description species: (Holten, 1802)
Genus: Hepatoxylon Bosc, 1811
Family: Sphyriocephalidae Pinter, 1913
Superfamily: Tentacularioidea
Suborder: Trypanobatoida
Order: Trypanorhyncha
Subclass: Eucestoda
Heterocotyle americana (1)
Hargis, 1955
  • Hypanus americanus (Hildebrand & Schroeder, 1928) (Myliobatiformes: Dasyatidae)
    • Chisholm, L.A. et.al. (1996) [9644] (used host: Dasyatis americana )
Original description species: Hargis, 1955
Genus: Heterocotyle Scott, 1904
Family: Monocotylidae Taschenberg, 1879
Order: Monocotylidea
Subclass: Monopisthocotylea
Heterocotyle armata (1)
Timofeeva, 1983
  • Himantura uarnak (Forsskål, 1775) (Myliobatiformes: Dasyatidae)
    • Chisholm, L.A. et.al. (1996) [9644]
Original description species: Timofeeva, 1983
Genus: Heterocotyle Scott, 1904
Family: Monocotylidae Taschenberg, 1879
Order: Monocotylidea
Subclass: Monopisthocotylea
Heterocotyle capapei (2)
Neifar, Euzet & Hassine, 2000
  • Dasyatis tortonesei Capapé, 1975 (Myliobatiformes: Dasyatidae)
    • Neifar, L. et.al. (2000) [7372]
    • Derbel, H. et.al. (2022) [31118]
Original description species: Neifar, Euzet & Hassine, 2000
Genus: Heterocotyle Scott, 1904
Family: Monocotylidae Taschenberg, 1879
Order: Monocotylidea
Subclass: Monopisthocotylea
Heterocotyle capricornensis (3)
Chisholm & Whittington, 1996
  • Pateobatis fai (Jordan & Seale, 1906) (Myliobatiformes: Dasyatidae)
    • Chisholm, L.A. et.al. (1996) [9644] (used host: Himantura fai )
    • Chisholm, L.A. (1998) [24082] (used host: Himantura fai )
    • Chisholm, L.A. et.al. (2001) [23992] (used host: Himantura fai )
Original description species: Chisholm & Whittington, 1996
Genus: Heterocotyle Scott, 1904
Family: Monocotylidae Taschenberg, 1879
Order: Monocotylidea
Subclass: Monopisthocotylea
Heterocotyle chinensis (2)
Timofeeva, 1983
  • Hemitrygon akajei (Müller & Henle, 1841) (Myliobatiformes: Dasyatidae)
    • Chisholm, L.A. et.al. (1996) [9644] (used host: Dasyatis akajei )
    • Nitta, M. et.al. (2015) [22929] (used host: Dasyatis akajei )
  • Hemitrygon fluviorum (Ogilby, 1908) (Myliobatiformes: Dasyatidae)
    • Chisholm, L.A. et.al. (1996) [9644] (used host: Dasyatis fluviorum )
  • Himantura uarnak (Forsskål, 1775) (Myliobatiformes: Dasyatidae)
    • Chisholm, L.A. et.al. (1996) [9644]
Original description species: Timofeeva, 1983
Genus: Heterocotyle Scott, 1904
Family: Monocotylidae Taschenberg, 1879
Order: Monocotylidea
Subclass: Monopisthocotylea
Heterocotyle confusa (1)
Timofeeva, 1983
  • Himantura uarnak (Forsskål, 1775) (Myliobatiformes: Dasyatidae)
    • Chisholm, L.A. et.al. (1996) [9644]
Original description species: Timofeeva, 1983
Genus: Heterocotyle Scott, 1904
Family: Monocotylidae Taschenberg, 1879
Order: Monocotylidea
Subclass: Monopisthocotylea
Heterocotyle dasyatis (1)
(Yamaguti, 1965)
Original description species: (Yamaguti, 1965)
Genus: Heterocotyle Scott, 1904
Family: Monocotylidae Taschenberg, 1879
Order: Monocotylidea
Subclass: Monopisthocotylea
Heterocotyle elliptica (1)
Pillai & Pillai, 1976
  • Pastinachus sephen (Fabricius in Niebuhr, 1775) (Myliobatiformes: Dasyatidae)
    • Chisholm, L.A. et.al. (1996) [9644]
Original description species: Pillai & Pillai, 1976
Genus: Heterocotyle Scott, 1904
Family: Monocotylidae Taschenberg, 1879
Order: Monocotylidea
Subclass: Monopisthocotylea
Heterocotyle forcifera (2)
Neifar, Euzet & Ben Hassine, 1999
  • Taeniurops grabatus (Geoffroy Saint-Hilaire, 1817) (Myliobatiformes: Dasyatidae)
    • Neifar, L. et.al. (1999) [21236] (used host: Taeniura grabata )
    • Derbel, H. et.al. (2022) [31118] (used host: Taeniura grabata )
Original description species: Neifar, Euzet & Ben Hassine, 1999
Genus: Heterocotyle Scott, 1904
Family: Monocotylidae Taschenberg, 1879
Order: Monocotylidea
Subclass: Monopisthocotylea
Heterocotyle granulatae (2)
Young, 1967
  • Himantura uarnak (Forsskål, 1775) (Myliobatiformes: Dasyatidae)
    • Chisholm, L.A. et.al. (1996) [9644]
  • Urogymnus granulatus (Macleay, 1883) (Myliobatiformes: Dasyatidae)
    • Young, P.C. (1967) [17104] (used host: Dasyatis granulata )
    • Chisholm, L.A. et.al. (1996) [9644] (used host: Himantura granulata )
Original description species: Young, 1967
Genus: Heterocotyle Scott, 1904
Family: Monocotylidae Taschenberg, 1879
Order: Monocotylidea
Subclass: Monopisthocotylea
Heterocotyle margaritae (3)
Chero, Cruces, Saez, Santos & Luque, 2020
  • Hypanus dipterurus (Jordan & Gilbert, 1880) (Myliobatiformes: Dasyatidae)
    • Chero, J.D. et.al. (2020) [28274]
    • Chero De La Cruz, J.D. (2022) [32535]
    • Santillán, L.A. et.al. (2024) [33202]
Original description species: Chero, Cruces, Saez, Santos & Luque, 2020
Genus: Heterocotyle Scott, 1904
Family: Monocotylidae Taschenberg, 1879
Order: Monocotylidea
Subclass: Monopisthocotylea
Heterocotyle minima (7)
(Maccallum, 1916)
  • Bathytoshia centroura (Mitchill, 1815) (Myliobatiformes: Dasyatidae)
    • Price, E.W. (1938) [17134] (used host: Pastinachus centrourus )
    • Sproston, N.G. (1946) [17150] (used host: Pastinachus centrourus )
    • Hendrix, S.S. (1994) [17205] (used host: Dasyatis centroura )
    • Chisholm, L.A. et.al. (1996) [9644] (used host: Dasyatis centroura )
    • Derbel, H. et.al. (2022) [31118]
  • Dasyatis pastinaca (Linnaeus, 1758) (Myliobatiformes: Dasyatidae)
    • Maccallum, G.A. (1916) [17158] (used host: Dasybatus pastinacus )
    • Price, E.W. (1938) [17134]
    • Sproston, N.G. (1946) [17150]
    • Young, P.C. (1967) [17104]
    • Chisholm, L.A. et.al. (1996) [9644]
  • Squalus acanthias Linnaeus, 1758 (Squaliformes: Squalidae)
Original description species: (Maccallum, 1916)
Genus: Heterocotyle Scott, 1904
Family: Monocotylidae Taschenberg, 1879
Order: Monocotylidea
Subclass: Monopisthocotylea
Heterocotyle mokhtarae (2)
Neifar, Euzet & Ben Hassine, 1999
  • Taeniurops grabatus (Geoffroy Saint-Hilaire, 1817) (Myliobatiformes: Dasyatidae)
    • Neifar, L. et.al. (1999) [21236] (used host: Taeniura grabata )
    • Derbel, H. et.al. (2022) [31118] (used host: Taeniura grabata )
Original description species: Neifar, Euzet & Ben Hassine, 1999
Genus: Heterocotyle Scott, 1904
Family: Monocotylidae Taschenberg, 1879
Order: Monocotylidea
Subclass: Monopisthocotylea
Heterocotyle pastinacae (7)
Scott, 1904
  • Dasyatis chrysonota (Smith, 1828) (Myliobatiformes: Dasyatidae)
    • Vaughan, D.B. et.al. (2024) [33260]
  • Dasyatis marmorata (Steindachner, 1892) (Myliobatiformes: Dasyatidae)
  • Dasyatis pastinaca (Linnaeus, 1758) (Myliobatiformes: Dasyatidae)
Original description species: Scott, 1904
Genus: Heterocotyle Scott, 1904
Family: Monocotylidae Taschenberg, 1879
Order: Monocotylidea
Subclass: Monopisthocotylea
Heterocotyle robusta (1)
(Johnston & Tiegs, 1922)
  • Trygonoptera testacea Müller & Henle, 1841 (Myliobatiformes: Urolophidae)
    • Chisholm, L.A. et.al. (1996) [9644]
Original description species: (Johnston & Tiegs, 1922)
Genus: Heterocotyle Scott, 1904
Family: Monocotylidae Taschenberg, 1879
Order: Monocotylidea
Subclass: Monopisthocotylea
Heterocotyle scotti (2)
Neifar, Euzet & Ben Hassine, 1998
  • Dasyatis marmorata (Steindachner, 1892) (Myliobatiformes: Dasyatidae)
  • Dasyatis pastinaca (Linnaeus, 1758) (Myliobatiformes: Dasyatidae)
Original description species: Neifar, Euzet & Ben Hassine, 1998
Genus: Heterocotyle Scott, 1904
Family: Monocotylidae Taschenberg, 1879
Order: Monocotylidea
Subclass: Monopisthocotylea
Heterocotyle similis (1)
Neifar, Euzet & Ben Hassine, 1998
  • Dasyatis pastinaca (Linnaeus, 1758) (Myliobatiformes: Dasyatidae)
Original description species: Neifar, Euzet & Ben Hassine, 1998
Genus: Heterocotyle Scott, 1904
Family: Monocotylidae Taschenberg, 1879
Order: Monocotylidea
Subclass: Monopisthocotylea
Heterocotyle sp. (3)

  • Pseudobatos planiceps (Garman, 1880) (Rhinopristiformes: Rhinobatidae)
    • Santillán, L.A. et.al. (2024) [33202]
  • Rhinoptera steindachneri Evermann & Jenkins, 1891 (Myliobatiformes: Rhinopteridae)
    • IVan Merlo-Serna, A. et.al. (2016) [23771]
    • Mendoza-Garfias, B. et.al. (2017) [26032]
Genus: Heterocotyle Scott, 1904
Family: Monocotylidae Taschenberg, 1879
Order: Monocotylidea
Subclass: Monopisthocotylea
Heterocotyle striata (2)
Neifar, Euzet & Ben Hassine, 1999
  • Taeniurops grabatus (Geoffroy Saint-Hilaire, 1817) (Myliobatiformes: Dasyatidae)
    • Neifar, L. et.al. (1999) [21236] (used host: Taeniura grabata )
    • Derbel, H. et.al. (2022) [31118] (used host: Taeniura grabata )
Original description species: Neifar, Euzet & Ben Hassine, 1999
Genus: Heterocotyle Scott, 1904
Family: Monocotylidae Taschenberg, 1879
Order: Monocotylidea
Subclass: Monopisthocotylea
Heterocotyle sulamericana (3)
Portes Santos, Santos, Cunha & Chisholm, 2012
  • Hypanus guttatus (Bloch & Schneider, 1801) (Myliobatiformes: Dasyatidae)
    • Santos, C.P. et.al. (2012) [14722] (used host: Dasyatis guttata )
    • Cohen, S.C. et.al. (2013) [22565] (used host: Dasyatis guttata )
    • Lopes, D.A. et.al. (2016) [24606] (used host: Dasyatis guttata )
Original description species: Portes Santos, Santos, Cunha & Chisholm, 2012
Genus: Heterocotyle Scott, 1904
Family: Monocotylidae Taschenberg, 1879
Order: Monocotylidea
Subclass: Monopisthocotylea
Heterocotyle taeniuropi (1)
Cao, Ding, Zhang & Liu, 2010
  • Taeniurops meyeni (Müller & Henle, 1841) (Myliobatiformes: Dasyatidae)
Original description species: Cao, Ding, Zhang & Liu, 2010
Genus: Heterocotyle Scott, 1904
Family: Monocotylidae Taschenberg, 1879
Order: Monocotylidea
Subclass: Monopisthocotylea
Heterocotyle tokoloshei (2)
Vaughan & Chisholm, 2010
  • Bathytoshia brevicaudata (Hutton, 1875) (Myliobatiformes: Dasyatidae)
    • Vaughan, D.B. et.al. (2010) [21201] (used host: Dasyatis brevicaudata )
    • Schaeffner, B.C. et.al. (2019) [27172]
Original description species: Vaughan & Chisholm, 2010
Genus: Heterocotyle Scott, 1904
Family: Monocotylidae Taschenberg, 1879
Order: Monocotylidea
Subclass: Monopisthocotylea
Heterocotyle whittingtoni (2)
Chisholm & Kritsky, 2020
  • Maculabatis toshi (Whitley, 1939) (Myliobatiformes: Dasyatidae)
    • Kritsky, D.C. et.al. (2020) [29167]
    • Chisholm, L.A. et.al. (2020) [29076]
Original description species: Chisholm & Kritsky, 2020
Genus: Heterocotyle Scott, 1904
Family: Monocotylidae Taschenberg, 1879
Order: Monocotylidea
Subclass: Monopisthocotylea
Heteronchocotyle gymnurae (3)
Neifar, Euzet & Ben Hassine, 2001
  • Gymnura altavela (Linnaeus, 1758) (Myliobatiformes: Gymnuridae)
  • Gymnura natalensis (Gilchrist & Thompson, 1911) (Myliobatiformes: Gymnuridae)
    • Vaughan, D.B. et.al. (2023) [31765]
Original description species: Neifar, Euzet & Ben Hassine, 2001
Genus: Heteronchocotyle Brooks, 1934
Family: Hexabothriidae Price, 1942
Order: Diclybothriidea
Subclass: Polyopisthocotylea
Heteronchocotyle hypoprioni (1)
Brooks, 1934
  • Negaprion brevirostris (Poey, 1868) (Carcharhiniformes: Carcharhinidae)
    • Sproston, N.G. (1946) [17150] (used host: Hypoprion brevirostris )
Original description species: Brooks, 1934
Genus: Heteronchocotyle Brooks, 1934
Family: Hexabothriidae Price, 1942
Order: Diclybothriidea
Subclass: Polyopisthocotylea
Heteronchocotyle leucas (3)
Hargis, 1955
  • Carcharhinus leucas (Valenciennes in Müller & Henle, 1839) (Carcharhiniformes: Carcharhinidae)
    • Hargis, W.J. (1955) [22560]
    • Watson, D.E. et.al. (1976) [16440]
    • Rodriguez-Ortiz, B. et.al. (2004) [16969]
Original description species: Hargis, 1955
Genus: Heteronchocotyle Brooks, 1934
Family: Hexabothriidae Price, 1942
Order: Diclybothriidea
Subclass: Polyopisthocotylea
Heteronybelinia australis (3)
Palm & Beveridge, 2002
  • Carcharhinus amboinensis (Müller & Henle, 1839) (Carcharhiniformes: Carcharhinidae)
  • Carcharhinus brachyurus (Günther, 1870) (Carcharhiniformes: Carcharhinidae)
  • Chiloscyllium punctatum Müller & Henle, 1838 (Orectolobiformes: Hemiscylliidae)
    • Schaeffner, B.C. et.al. (2014) [21816] ( used parasite: Heteronybelinia cf. australis)
Synonyms: Heteronybelinia cf. australis
Original description species: Palm & Beveridge, 2002
Genus: Heteronybelinia Palm, 1999
Family: Tentaculariidae Poche, 1926
Superfamily: Tentacularioidea
Suborder: Trypanobatoida
Order: Trypanorhyncha
Subclass: Eucestoda
Heteronybelinia estigmena (18)
(Dollfus, 1960)
  • Beringraja binoculata (Girard, 1855) (Rajiformes: Rajidae)
    • Palm, H.W. (2004) [16112] (used host: Raja binoculata )
  • Caliraja inornata (Jordan & Gilbert, 1881) (Rajiformes: Rajidae)
    • Palm, H.W. (2004) [16112] (used host: Raja inornata )
  • Caliraja rhina (Jordan & Gilbert, 1880) (Rajiformes: Rajidae)
    • Palm, H.W. (2004) [16112] (used host: Raja rhina )
  • Carcharhinus amblyrhynchoides (Whitley, 1934) (Carcharhiniformes: Carcharhinidae)
  • Carcharhinus brevipinna (Müller & Henle, 1839) (Carcharhiniformes: Carcharhinidae)
    • Beveridge, I. et.al. (2014) [21718]
  • Carcharhinus leucas (Valenciennes in Müller & Henle, 1839) (Carcharhiniformes: Carcharhinidae)
    • Watson, D.E. et.al. (1976) [16440] ( used parasite: Nybelinia bisulcata)
    • Palm, H.W. (1999) [16366]
    • Rodriguez-Ortiz, B. et.al. (2004) [16969] ( used parasite: Nybelinia bisulcata)
    • Palm, H.W. (2004) [16112]
    • Cutmore, S.C. et.al. (2009) [7421]
    • Schaeffner, B.C. et.al. (2014) [21816]
    • Schaeffner, B.C. et.al. (2019) [27172]
    • Palm, H.W. et.al. (2023) [32432] (used host: Carcharhinus cf. leucas )
  • Carcharhinus limbatus (Valenciennes in Müller & Henle, 1839) (Carcharhiniformes: Carcharhinidae)
    • Palm, H.W. (1995) [17852] ( used parasite: Nybelinia alloiotica mihi)
    • Palm, H.W. (1999) [16366]
    • Palm, H.W. et.al. (2000) [16209] ( used parasite: Nybelinia cf. bisulcata)
    • Palm, H.W. et.al. (2000) [16209]
    • Palm, H.W. et.al. (2002) [17845]
    • Palm, H.W. (2004) [16112] ( used parasite: Nybelinia cf. bisulcata)
    • Palm, H.W. (2004) [16112]
    • Palm, H.W. et.al. (2009) [15747] ( used parasite: Heteronybelinia cf. estigmena )
    • Schaeffner, B.C. et.al. (2014) [21816]
    • Schaeffner, B.C. et.al. (2019) [27172]
    • Palm, H.W. et.al. (2023) [32432] (used host: Carcharhinus cf. limbatus )
  • Carcharhinus obscurus (Lesueur, 1818) (Carcharhiniformes: Carcharhinidae)
    • Linton, E. (1905) [17126] ( used parasite: Tetrarhynchus bisulcatus)
    • Palm, H.W. (2004) [16112]
  • Carcharhinus plumbeus (Nardo, 1827) (Carcharhiniformes: Carcharhinidae)
    • Linton, E. (1905) [17126] (used host: Carcharhinus milberti | used parasite: Tetrarhynchus bisulcatus)
  • Notorynchus cepedianus (Péron, 1807) (Hexanchiformes: Hexanchidae)
    • de São Clemente, S.C. et.al. (1992) [17922] (used host: Notorhynchus pectorosus | used parasite: Nybelinia bisulcata)
  • Prionace glauca (Linnaeus, 1758) (Carcharhiniformes: Carcharhinidae)
  • Rhizoprionodon terraenovae (Richardson, 1836) (Carcharhiniformes: Carcharhinidae)
    • Linton, E. (1905) [17126] (used host: Scoliodon terrae-novae | used parasite: Tetrarhynchus bisulcatus)
  • Sphyrna zygaena (Linnaeus, 1758) (Carcharhiniformes: Sphyrnidae)
    • Maccallum, G.A. (1921) [16352] (used host: Cestracion zygaena | used parasite: Tetrarhynchus bisulcatum)
Synonyms: Heteronybelinia cf. estigmena , Nybelinia alloiotica mihi, Nybelinia bisulcata, Nybelinia cf. bisulcata, Tetrarhynchus bisulcatum, Tetrarhynchus bisulcatus
Original description species: (Dollfus, 1960)
Genus: Heteronybelinia Palm, 1999
Family: Tentaculariidae Poche, 1926
Superfamily: Tentacularioidea
Suborder: Trypanobatoida
Order: Trypanorhyncha
Subclass: Eucestoda
Heteronybelinia eureia (2)
(Dollfus, 1960)
  • Mustelus canis (Mitchill, 1815) (Carcharhiniformes: Triakidae)
Original description species: (Dollfus, 1960)
Genus: Heteronybelinia Palm, 1999
Family: Tentaculariidae Poche, 1926
Superfamily: Tentacularioidea
Suborder: Trypanobatoida
Order: Trypanorhyncha
Subclass: Eucestoda
Heteronybelinia heteromorphi (3)
Palm, 1999
  • Eusphyra blochii (Cuvier, 1816) (Carcharhiniformes: Sphyrnidae)
    • Palm, H.W. (1999) [16366] (used host: Sphyrna blochii )
    • Palm, H.W. (2004) [16112]
  • Sphyrna mokarran (Rüppell, 1837) (Carcharhiniformes: Sphyrnidae)
Original description species: Palm, 1999
Genus: Heteronybelinia Palm, 1999
Family: Tentaculariidae Poche, 1926
Superfamily: Tentacularioidea
Suborder: Trypanobatoida
Order: Trypanorhyncha
Subclass: Eucestoda
Heteronybelinia mattisi (4)
Menoret & Ivanov, 2013
  • Sympterygia bonapartii Müller & Henle, 1841 (Rajiformes: Arhynchobatidae)
    • Menoret, A. et.al. (2013) [20541]
    • Irigoitia, M.M. et.al. (2017) [25811]
    • Alves, P.V. et.al. (2017) [25154]
    • Franzese, S. et.al. (2023) [31906]
Original description species: Menoret & Ivanov, 2013
Genus: Heteronybelinia Palm, 1999
Family: Tentaculariidae Poche, 1926
Superfamily: Tentacularioidea
Suborder: Trypanobatoida
Order: Trypanorhyncha
Subclass: Eucestoda
Heteronybelinia nipponica (5)
(Yamaguti, 1952)
  • Carcharhinus signatus (Poey, 1868) (Carcharhiniformes: Carcharhinidae)
    • Muniz-Pereira, L.C. et.al. (2009) [21220]
    • Alves, P.V. et.al. (2017) [25154]
  • Rhynchobatus djiddensis (Forsskål, 1775) (Rhinopristiformes: Rhinidae)
  • Sphyrna lewini (Griffith & Smith, 1834) (Carcharhiniformes: Sphyrnidae)
    • de São Clemente, S.C. et.al. (1992) [17922] ( used parasite: Nybelinia rougetcampanae)
    • Palm, H.W. (2004) [16112]
    • Alves, P.V. et.al. (2017) [25154]
  • Sphyrna zygaena (Linnaeus, 1758) (Carcharhiniformes: Sphyrnidae)
    • Gómes, D.C. et.al. (2005) [10843] ( used parasite: Heteronybelinia rougetcampanae)
    • Alves, P.V. et.al. (2017) [25154]
Synonyms: Heteronybelinia rougetcampanae, Nybelinia nipponica, Nybelinia rougetcampanae
Original description species: (Yamaguti, 1952)
Genus: Heteronybelinia Palm, 1999
Family: Tentaculariidae Poche, 1926
Superfamily: Tentacularioidea
Suborder: Trypanobatoida
Order: Trypanorhyncha
Subclass: Eucestoda
Heteronybelinia overstreeti (1)
Palm, 2004
  • Carcharhinus limbatus (Valenciennes in Müller & Henle, 1839) (Carcharhiniformes: Carcharhinidae)
Original description species: Palm, 2004
Genus: Heteronybelinia Palm, 1999
Family: Tentaculariidae Poche, 1926
Superfamily: Tentacularioidea
Suborder: Trypanobatoida
Order: Trypanorhyncha
Subclass: Eucestoda
Heteronybelinia palliata (3)
(Linton, 1924)
  • Mustelus canis (Mitchill, 1815) (Carcharhiniformes: Triakidae)
  • Notorynchus cepedianus (Péron, 1807) (Hexanchiformes: Hexanchidae)
  • Sphyrna zygaena (Linnaeus, 1758) (Carcharhiniformes: Sphyrnidae)
    • Chandler, A.C. (1942) [16290] ( used parasite: Nybelinia palliata)
    • Palm, H.W. et.al. (2000) [16209]
    • Palm, H.W. (2004) [16112]
Synonyms: Nybelinia palliata
Original description species: (Linton, 1924)
Genus: Heteronybelinia Palm, 1999
Family: Tentaculariidae Poche, 1926
Superfamily: Tentacularioidea
Suborder: Trypanobatoida
Order: Trypanorhyncha
Subclass: Eucestoda
Heteronybelinia perideraeus (4)
(Shipley & Hornell, 1906)
  • Glyphis gangeticus (Müller & Henle, 1839) (Carcharhiniformes: Carcharhinidae)
    • Palm, H.W. et.al. (1999) [17850] (used host: Carcharhinus gangeticus | used parasite: Nybelinia perideraeus)
    • Palm, H.W. (2004) [16112]
  • Nebrius ferrugineus (Lesson, 1831) (Orectolobiformes: Ginglymostomatidae)
  • Notorynchus cepedianus (Péron, 1807) (Hexanchiformes: Hexanchidae)
  • Rhynchobatus djiddensis (Forsskål, 1775) (Rhinopristiformes: Rhinidae)
  • Scoliodon laticaudus Müller & Henle, 1838 (Carcharhiniformes: Carcharhinidae)
    • Vijayalakshmi, C. et.al. (1996) [17851] (used host: Scoliodon palasorrah | used parasite: Nybelinia perideraeus)
Synonyms: Nybelinia perideraeus, Tetrarhynchus perideraeus
Original description species: (Shipley & Hornell, 1906)
Genus: Heteronybelinia Palm, 1999
Family: Tentaculariidae Poche, 1926
Superfamily: Tentacularioidea
Suborder: Trypanobatoida
Order: Trypanorhyncha
Subclass: Eucestoda
Heteronybelinia robusta (5)
(Linton, 1890)
  • Bathytoshia centroura (Mitchill, 1815) (Myliobatiformes: Dasyatidae)
    • Palm, H.W. (1999) [16366] (used host: Dasyatis centroura )
    • Palm, H.W. (2004) [16112] (used host: Dasyatis centroura )
  • Caliraja inornata (Jordan & Gilbert, 1881) (Rajiformes: Rajidae)
    • Palm, H.W. (2004) [16112] (used host: Raja inornata )
  • Carcharhinus limbatus (Valenciennes in Müller & Henle, 1839) (Carcharhiniformes: Carcharhinidae)
  • Mustelus asterias Cloquet, 1819 (Carcharhiniformes: Triakidae)
  • Rajella leoparda (Von Bonde & Swart, 1923) (Rajiformes: Rajidae)
    • Palm, H.W. (2004) [16112] (used host: Rajella leopardus )
Original description species: (Linton, 1890)
Genus: Heteronybelinia Palm, 1999
Family: Tentaculariidae Poche, 1926
Superfamily: Tentacularioidea
Suborder: Trypanobatoida
Order: Trypanorhyncha
Subclass: Eucestoda
Heteronybelinia sp. (2)

  • Caliraja inornata (Jordan & Gilbert, 1881) (Rajiformes: Rajidae)
    • Palm, H.W. (2004) [16112] (used host: Raja inornata )
  • Tetronarce nobiliana (Bonaparte, 1835) (Torpediniformes: Torpedinidae)
    • Palm, H.W. (2004) [16112] (used host: Torpedo nobiliana )
  • Zearaja brevicaudata (Marini, 1933) (Rajiformes: Rajidae)
    • Irigoitia, M.M. et.al. (2022) [31017] (used host: Dipturus brevicaudata )
Genus: Heteronybelinia Palm, 1999
Family: Tentaculariidae Poche, 1926
Superfamily: Tentacularioidea
Suborder: Trypanobatoida
Order: Trypanorhyncha
Subclass: Eucestoda
Heteronybelinia yamagutii (5)
(Dollfus, 1960)
  • Carcharhinus signatus (Poey, 1868) (Carcharhiniformes: Carcharhinidae)
    • Muniz-Pereira, L.C. et.al. (2009) [21220]
    • Alves, P.V. et.al. (2017) [25154]
  • Sphyrna lewini (Griffith & Smith, 1834) (Carcharhiniformes: Sphyrnidae)
Original description species: (Dollfus, 1960)
Genus: Heteronybelinia Palm, 1999
Family: Tentaculariidae Poche, 1926
Superfamily: Tentacularioidea
Suborder: Trypanobatoida
Order: Trypanorhyncha
Subclass: Eucestoda
Heterosphyriocephalus encarnae (1)
Dallares, Carrasson & Schaeffner, 2017
  • Alopias pelagicus Nakamura, 1935 (Lamniformes: Alopiidae)
    • Dallares, S. et.al. (2017) [25091]
Original description species: Dallares, Carrasson & Schaeffner, 2017
Genus: Heterosphyriocephalus Palm, 2004
Family: Sphyriocephalidae Pinter, 1913
Superfamily: Tentacularioidea
Suborder: Trypanobatoida
Order: Trypanorhyncha
Subclass: Eucestoda
Heterosphyriocephalus tergestinus (3)
(Pintner, 1913)
  • Alopias vulpinus (Bonnaterre, 1788) (Lamniformes: Alopiidae)
    • Palm, H.W. (2004) [16112] ( used parasite: Sphyriocephalus tergestinus)
    • Dallares, S. et.al. (2017) [25091]
  • Etmopterus spinax (Linnaeus, 1758) (Squaliformes: Etmopteridae)
    • Santoro, M. et.al. (2022) [31384]
  • Euprotomicrus bispinatus (Quoy & Gaimard, 1824) (Squaliformes: Dalatiidae)
    • Palm, H.W. (2004) [16112] ( used parasite: Sphyriocephalus tergestinus)
    • Dallares, S. et.al. (2017) [25091]
  • Isurus oxyrinchus Rafinesque, 1810 (Lamniformes: Lamnidae)
    • Palm, H.W. (2004) [16112] ( used parasite: Sphyriocephalus tergestinus)
    • Dallares, S. et.al. (2017) [25091]
  • Prionace glauca (Linnaeus, 1758) (Carcharhiniformes: Carcharhinidae)
    • Dallares, S. et.al. (2017) [25091] (used host: Isurus glaucus )
Synonyms: Sphyriocephalus tergestinus
Original description species: (Pintner, 1913)
Genus: Heterosphyriocephalus Palm, 2004
Family: Sphyriocephalidae Pinter, 1913
Superfamily: Tentacularioidea
Suborder: Trypanobatoida
Order: Trypanorhyncha
Subclass: Eucestoda
Hexabothrium akaroensis (2)
Dillon & Hargis, 1965
  • Galeorhinus galeus (Linnaeus, 1758) (Carcharhiniformes: Triakidae)
    • Dillon, W.A. et.al. (1968) [22557] (used host: Galeorhinus australis )
    • Hewitt, G.C. et.al. (1972) [16602] (used host: Galeorhinus australis )
Original description species: Dillon & Hargis, 1965
Genus: Hexabothrium Von Nordmann, 1832
Family: Hexabothriidae Price, 1942
Order: Diclybothriidea
Subclass: Polyopisthocotylea
Hexabothrium appendiculatum (13)
(Kuhn, 1829)
  • Dipturus batis (Linnaeus, 1758) (Rajiformes: Rajidae)
    • Olsson, P. (1876) [22616] (used host: Raja batis | used parasite: Onchocotyle appendiculata)
    • Baylis, H.A. et.al. (1933) [19885] (used host: Raja batis | used parasite: Onchocotyle appendiculata)
  • Dipturus oxyrinchus (Linnaeus, 1758) (Rajiformes: Rajidae)
    • Baylis, H.A. et.al. (1933) [19885] (used host: Raja oxyrhynchus | used parasite: Onchocotyle appendiculata)
  • Raja brachyura Lafont, 1873 (Rajiformes: Rajidae)
    • Baylis, H.A. et.al. (1933) [19885] (used host: Raja blanda | used parasite: Onchocotyle appendiculata)
  • Raja clavata Linnaeus, 1758 (Rajiformes: Rajidae)
    • Baylis, H.A. et.al. (1933) [19885] ( used parasite: Onchocotyle appendiculata)
  • Scyliorhinus canicula (Linnaeus, 1758) (Carcharhiniformes: Scyliorhinidae)
    • Baylis, H.A. et.al. (1933) [19885] (used host: Scyllium canicula | used parasite: Onchocotyle appendiculata)
    • Llewellyn, J. et.al. (1984) [15273]
    • Whittington, I.D. (1987) [17450]
    • Whittington, I.D. (1987) [17449]
    • Whittington, I.D. (1987) [17451]
    • Mollaret, I. et.al. (2000) [21218]
    • Moore, A.B.M. (2001) [17250]
    • Dallares, S. et.al. (2017) [25639]
  • Scyliorhinus stellaris (Linnaeus, 1758) (Carcharhiniformes: Scyliorhinidae)
    • Cerfontaine, P. (1899) [22589] (used host: Scyllium catulus | used parasite: Acanthonchocotyle appendiculata)
    • Sproston, N.G. (1946) [17150] (used host: Scylliorhinus catulus )
    • Llewellyn, J. et.al. (1984) [15273]
    • Derbel, H. et.al. (2022) [31118]
    • Santoro, M. et.al. (2023) [31341]
Synonyms: Acanthonchocotyle appendiculata, Onchocotyle appendiculata
Original description species: (Kuhn, 1829)
Genus: Hexabothrium Von Nordmann, 1832
Family: Hexabothriidae Price, 1942
Order: Diclybothriidea
Subclass: Polyopisthocotylea
Hexabothrium canicula (3)
(Cerfontaine, 1899)
  • Scyliorhinus canicula (Linnaeus, 1758) (Carcharhiniformes: Scyliorhinidae)
    • Cerfontaine, P. (1899) [22589] (used host: Scyllium canicula | used parasite: Acanthonchocotyle canicula)
    • Sproston, N.G. (1946) [17150]
    • Henderson, A.C. et.al. (1998) [9344]
Synonyms: Acanthonchocotyle canicula
Original description species: (Cerfontaine, 1899)
Genus: Hexabothrium Von Nordmann, 1832
Family: Hexabothriidae Price, 1942
Order: Diclybothriidea
Subclass: Polyopisthocotylea
Hexabothrium musteli (4)
(Maccallum, 1931) Price, 1942
  • Mustelus canis (Mitchill, 1815) (Carcharhiniformes: Triakidae)
    • Maccallum, G.A. (1931) [22561] ( used parasite: Acanthonchocotyle musteli)
    • Hendrix, S.S. (1994) [17205]
  • Mustelus mustelus (Linnaeus, 1758) (Carcharhiniformes: Triakidae)
Synonyms: Acanthonchocotyle musteli
Original description species: (Maccallum, 1931) Price, 1942
Genus: Hexabothrium Von Nordmann, 1832
Family: Hexabothriidae Price, 1942
Order: Diclybothriidea
Subclass: Polyopisthocotylea
Hexacanalis abruptus (4)
(Southwell, 1911) Perrenoud, 1931
  • Gymnura micrura (Bloch & Schneider, 1801) (Myliobatiformes: Gymnuridae)
    • Southwell, T. (1911) [17129] (used host: Pteroplatea micrura | used parasite: Cephalobothrium abruptum)
    • Jensen, K. (2005) [16338]
    • Cielocha, J.J. et.al. (2011) [13082]
  • Neotrygon kuhlii (Müller & Henle, 1841) (Myliobatiformes: Dasyatidae)
    • Southwell, T. (1927) [16410] (used host: Trygon kuhlii | used parasite: Cephalobothrium abruptum)
Synonyms: Cephalobothrium abruptum
Original description species: (Southwell, 1911) Perrenoud, 1931
Genus: Hexacanalis Perrenoud, 1931
Family: Lecanicephalidae Braun, 1900
Order: Lecanicephalidea
Subclass: Eucestoda
Hexacanalis folifer (3)
Cielocha & Jensen, 2011
  • Gymnura zonura (Bleeker, 1852) (Myliobatiformes: Gymnuridae)
    • Cielocha, J.J. et.al. (2011) [13082]
    • Borucinska, J.D. et.al. (2013) [17367]
    • Jensen, K. et.al. (2016) [23959]
Original description species: Cielocha & Jensen, 2011
Genus: Hexacanalis Perrenoud, 1931
Family: Lecanicephalidae Braun, 1900
Order: Lecanicephalidea
Subclass: Eucestoda
Hexacanalis govindi (1)
Wankhede, 2003
  • Mobula eregoodoo (Cantor, 1849) (Myliobatiformes: Mobulidae)
    • Wankhede, H. (2003) [16438] (used host: Dicerobatis eregoodoo )
Original description species: Wankhede, 2003
Genus: Hexacanalis Perrenoud, 1931
Family: Lecanicephalidae Braun, 1900
Order: Lecanicephalidea
Subclass: Eucestoda
Hexacanalis indirajii (1)
Murlidhar, 1986
  • Pastinachus sephen (Fabricius in Niebuhr, 1775) (Myliobatiformes: Dasyatidae)
    • Murlidhar, A. (1986) [16362] (used host: Trygon sephen )
Original description species: Murlidhar, 1986
Genus: Hexacanalis Perrenoud, 1931
Family: Lecanicephalidae Braun, 1900
Order: Lecanicephalidea
Subclass: Eucestoda
Hexacanalis pteroplateae (2)
(Zaidi & Khan, 1976) Cielocha & Jensen, 2011
  • Gymnura micrura (Bloch & Schneider, 1801) (Myliobatiformes: Gymnuridae)
    • Zaidi, D.A. et.al. (1976) [16468] (used host: Pteroplatea micrura | used parasite: Cephalobothrium pteroplateai)
    • Cielocha, J.J. et.al. (2011) [13082]
Synonyms: Cephalobothrium pteroplateai
Original description species: (Zaidi & Khan, 1976) Cielocha & Jensen, 2011
Genus: Hexacanalis Perrenoud, 1931
Family: Lecanicephalidae Braun, 1900
Order: Lecanicephalidea
Subclass: Eucestoda
Hexacanalis sasoonensis (1)
Srivastava & Capoor, 1980
  • Himantura marginata (Blyth, 1860) (Myliobatiformes: Dasyatidae)
    • Srivastav, A.K. et.al. (1980) [16422] (used host: Trygon marginatus )
Original description species: Srivastava & Capoor, 1980
Genus: Hexacanalis Perrenoud, 1931
Family: Lecanicephalidae Braun, 1900
Order: Lecanicephalidea
Subclass: Eucestoda
Hexacanalis smythii (1)
Murlidhar, Shinde & Jadhav, 1986
  • Bathytoshia centroura (Mitchill, 1815) (Myliobatiformes: Dasyatidae)
    • Murlidhar, A. et.al. (1986) [16363] (used host: Trygon centrura )
Original description species: Murlidhar, Shinde & Jadhav, 1986
Genus: Hexacanalis Perrenoud, 1931
Family: Lecanicephalidae Braun, 1900
Order: Lecanicephalidea
Subclass: Eucestoda
Hexacanalis thapari (2)
Shinde, Jadhav & Jadhav, 1991
  • Telatrygon zugei (Bürger in Müller & Henle, 1841) (Myliobatiformes: Dasyatidae)
    • Shinde, G.B. et.al. (1991) [16404] (used host: Trygon zugei )
    • Mohekar, A.D. et.al. (2002) [16360] (used host: Trygon zugei )
Original description species: Shinde, Jadhav & Jadhav, 1991
Genus: Hexacanalis Perrenoud, 1931
Family: Lecanicephalidae Braun, 1900
Order: Lecanicephalidea
Subclass: Eucestoda
Hexacanalis yamagutii (1)
Shinde & Deshmukh, 1979
  • Mobula eregoodoo (Cantor, 1849) (Myliobatiformes: Mobulidae)
    • Shinde, G.B. et.al. (1979) [16399] (used host: Dicerobatis eregoodoo )
Original description species: Shinde & Deshmukh, 1979
Genus: Hexacanalis Perrenoud, 1931
Family: Lecanicephalidae Braun, 1900
Order: Lecanicephalidea
Subclass: Eucestoda
Hexacanalis zugeis (1)
Shinde & Deshmukh, 1979
  • Telatrygon zugei (Bürger in Müller & Henle, 1841) (Myliobatiformes: Dasyatidae)
    • Shinde, G.B. et.al. (1979) [16399] (used host: Trygon zugei )
Original description species: Shinde & Deshmukh, 1979
Genus: Hexacanalis Perrenoud, 1931
Family: Lecanicephalidae Braun, 1900
Order: Lecanicephalidea
Subclass: Eucestoda
Hispidorhynchus styracurae (1)
Schaeffner, 2018
  • Styracura schmardae (Werner, 1904) (Myliobatiformes: Potamotrygonidae)
Original description species: Schaeffner, 2018
Genus: Hispidorhynchus Schaeffner & Beveridge, 2012
Family: Eutetrarhynchidae Guiart, 1927
Superfamily: Eutetrarhynchoidea
Suborder: Trypanobatoida
Order: Trypanorhyncha
Subclass: Eucestoda
Histodytes microocellatus (2)
Aragort, Alvarez, Iglesias, Leiro & Sanmartín, 2002
  • Raja microocellata Montagu, 1818 (Rajiformes: Rajidae)
    • Aragort, W. et.al. (2002) [10848]
    • Alvarez, M.F. et.al. (2006) [9086]
Original description species: Aragort, Alvarez, Iglesias, Leiro & Sanmartín, 2002
Genus: Histodytes Aragort, Alvarez, Iglesias, Leiro & Sanmartin, 2002
Family: Guyanemidae Petter, 1975
Superfamily: Dracunculoidea
Suborder: Camallanina
Order: Spirurida
Holobomolochus sp. (1)

  • Raja brachyura Lafont, 1873 (Rajiformes: Rajidae)
    • Alvarez, M.F. et.al. (2006) [9086]
  • Raja microocellata Montagu, 1818 (Rajiformes: Rajidae)
    • Alvarez, M.F. et.al. (2006) [9086]
  • Raja undulata Lacepède, 1802 (Rajiformes: Rajidae)
    • Alvarez, M.F. et.al. (2006) [9086]
Genus: Holobomolochus Vervoort, 1969
Family: Bomolochidae Claus, 1875
Order: Poecilostomatoida
Superorder: Podoplea
Subclass: Copepoda
Holocephalocotyle monstrosae (1)
Derouiche, Neifar, Gey, Justine & Tazerouti, 2019
  • Chimaera monstrosa Linnaeus, 1758 (Chimaeriformes: Chimaeridae)
    • Derouiche, I. et.al. (2019) [27591]
Original description species: Derouiche, Neifar, Gey, Justine & Tazerouti, 2019
Genus: Holocephalocotyle Derouiche, Neifar, Gey, Justine & Tazerouti, 2019
Family: Monocotylidae Taschenberg, 1879
Order: Monocotylidea
Subclass: Monopisthocotylea
Homeokotorella anterioporus (1)
Palm & Haseli, 2023
  • Taeniura lymma (Fabricius in Niebuhr, 1775) (Myliobatiformes: Dasyatidae)
Original description species: Palm & Haseli, 2023
Genus: Homeokotorella Palm & Haseli, 2023
Family: Tentaculariidae Poche, 1926
Superfamily: Tentacularioidea
Suborder: Trypanobatoida
Order: Trypanorhyncha
Subclass: Eucestoda
Homestios janinecairae (1)
Warren & Bullard, 2021
  • Aetomylaeus nichofii (Bloch & Schneider, 1801) (Myliobatiformes: Myliobatidae)
    • Warren, M.B. et.al. (2021) [29951]
Original description species: Warren & Bullard, 2021
Genus: Homestios Warren & Bullard, 2021
Family: Aporocotylidae Odhner, 1912
Superfamily: Schistosomatoidea
Suborder: Diplostomata
Order: Diplostomida
Subclass: Digenea
Hornelliella annandalei (5)
(Shipley & Hornell, 1912) Yamaguti, 1954
  • Chiloscyllium punctatum Müller & Henle, 1838 (Orectolobiformes: Hemiscylliidae)
    • Campbell, R.A. et.al. (1987) [16278] ( used parasite: Hornelliella macropora)
    • Palm, H.W. (2004) [16112]
  • Galeocerdo cuvier (Péron & Lesueur, 1822) (Carcharhiniformes: Galeocerdonidae)
  • Himantura uarnak (Forsskål, 1775) (Myliobatiformes: Dasyatidae)
  • Stegostoma tigrinum (Forster, 1781) (Orectolobiformes: Stegostomatidae)
    • Yamaguti, S. (1954) [16460]
    • Campbell, R.A. et.al. (1987) [16278] (used host: Stegostoma fasciatum | used parasite: Hornelliella macropora)
    • Palm, H.W. (2004) [16112] (used host: Stegostoma fasciatum )
    • Palm, H.W. et.al. (2009) [15747] (used host: Stegostoma fasciatum )
    • Schaeffner, B.C. et.al. (2014) [21816] (used host: Stegostoma fasciatum )
Synonyms: Hornelliella macropora
Original description species: (Shipley & Hornell, 1912) Yamaguti, 1954
Genus: Hornelliella Yamaguti, 1954
Family: Lacistorhynchidae Guiart, 1927
Superfamily: Lacistorhynchoidea
Suborder: Trypanoselachoida
Order: Trypanorhyncha
Subclass: Eucestoda
Hornelliella palasoorahi (2)
Zaidi & Khan, 1976
  • Rhizoprionodon acutus (Rüppell, 1837) (Carcharhiniformes: Carcharhinidae)
  • Scoliodon laticaudus Müller & Henle, 1838 (Carcharhiniformes: Carcharhinidae)
    • Zaidi, D.A. et.al. (1976) [16468] (used host: Scoliodon palasorrah )
Original description species: Zaidi & Khan, 1976
Genus: Hornelliella Yamaguti, 1954
Family: Lacistorhynchidae Guiart, 1927
Superfamily: Lacistorhynchoidea
Suborder: Trypanoselachoida
Order: Trypanorhyncha
Subclass: Eucestoda
Hornellobothrium cobraformis (2)
Shipley & Hornell, 1906
  • Aetobatus narinari (Euphrasen, 1790) (Myliobatiformes: Aetobatidae)
Original description species: Shipley & Hornell, 1906
Genus: Hornellobothrium Shipley & Hornell, 1906
Family: Lecanicephalidea incertae sedis Braun, 1900
Order: Lecanicephalidea
Subclass: Eucestoda
Hornellobothrium extensivum (1)
Jensen, 2005
  • Aetobatus narinari (Euphrasen, 1790) (Myliobatiformes: Aetobatidae)
Original description species: Jensen, 2005
Genus: Hornellobothrium Shipley & Hornell, 1906
Family: Lecanicephalidea incertae sedis Braun, 1900
Order: Lecanicephalidea
Subclass: Eucestoda
Hornellobothrium gerdaae (1)
Mojica, Jensen & Caira, 2014
  • Aetobatus ocellatus (Kuhl, 1823) (Myliobatiformes: Aetobatidae)
    • Mojica, K.R. et.al. (2014) [21567]
Original description species: Mojica, Jensen & Caira, 2014
Genus: Hornellobothrium Shipley & Hornell, 1906
Family: Lecanicephalidea incertae sedis Braun, 1900
Order: Lecanicephalidea
Subclass: Eucestoda
Hornellobothrium iotakotta (1)
Mojica, Jensen & Caira, 2014
  • Aetobatus ocellatus (Kuhl, 1823) (Myliobatiformes: Aetobatidae)
    • Mojica, K.R. et.al. (2014) [21567]
Original description species: Mojica, Jensen & Caira, 2014
Genus: Hornellobothrium Shipley & Hornell, 1906
Family: Lecanicephalidea incertae sedis Braun, 1900
Order: Lecanicephalidea
Subclass: Eucestoda
Hornellobothrium kolossakotta (1)
Mojica, Jensen & Caira, 2014
  • Aetobatus ocellatus (Kuhl, 1823) (Myliobatiformes: Aetobatidae)
    • Mojica, K.R. et.al. (2014) [21567]
Original description species: Mojica, Jensen & Caira, 2014
Genus: Hornellobothrium Shipley & Hornell, 1906
Family: Lecanicephalidea incertae sedis Braun, 1900
Order: Lecanicephalidea
Subclass: Eucestoda
Hornellobothrium najaforme (2)
Mojica, Jensen & Caira, 2014
  • Aetobatus ocellatus (Kuhl, 1823) (Myliobatiformes: Aetobatidae)
Original description species: Mojica, Jensen & Caira, 2014
Genus: Hornellobothrium Shipley & Hornell, 1906
Family: Lecanicephalidea incertae sedis Braun, 1900
Order: Lecanicephalidea
Subclass: Eucestoda
Hornellobothrium sp. (1)

  • Aetobatus ocellatus (Kuhl, 1823) (Myliobatiformes: Aetobatidae)
Genus: Hornellobothrium Shipley & Hornell, 1906
Family: Lecanicephalidea incertae sedis Braun, 1900
Order: Lecanicephalidea
Subclass: Eucestoda
Huffmanela carcharhini (3)
(Maccallum, 1925)
  • Carcharhinus melanopterus (Quoy & Gaimard, 1824) (Carcharhiniformes: Carcharhinidae)
    • Pereira, F.B. et.al. (2023) [31368]
  • Carcharhinus obscurus (Lesueur, 1818) (Carcharhiniformes: Carcharhinidae)
    • Maccallum, G.A. (1925) [16959] (used host: Carcharhinus commersoni | used parasite: Capillaria carcharhini)
  • Carcharhinus plumbeus (Nardo, 1827) (Carcharhiniformes: Carcharhinidae)
    • Diouf, J.N. et.al. (2003) [15715] ( used parasite: Huffmanela cf. carcharhini)
Synonyms: Capillaria carcharhini, Huffmanela cf. carcharhini
Original description species: (Maccallum, 1925)
Genus: Huffmanela Moravec, 1987
Family: Trichosomoididae Yorke & Maplestone, 1926
Huffmanela lata (1)
Justine, 2005
  • Carcharhinus amblyrhynchos (Bleeker, 1856) (Carcharhiniformes: Carcharhinidae)
    • Justine, J.-L. (2005) [7397]
Original description species: Justine, 2005
Genus: Huffmanela Moravec, 1987
Family: Trichosomoididae Yorke & Maplestone, 1926
Huffmanela markgracei (2)
Ruiz & Bullard, 2013
  • Rhizoprionodon terraenovae (Richardson, 1836) (Carcharhiniformes: Carcharhinidae)
    • Ruiz, C.F. et.al. (2013) [19638]
    • Pereira, F.B. et.al. (2023) [31368]
Original description species: Ruiz & Bullard, 2013
Genus: Huffmanela Moravec, 1987
Family: Trichosomoididae Yorke & Maplestone, 1926
Huffmanela selachii (1)
Al-Sabi, Ibrahim,Al-Hizab, Al-Jabr, Al-Shubaythi & Huffman, 2022
  • Carcharhinus melanopterus (Quoy & Gaimard, 1824) (Carcharhiniformes: Carcharhinidae)
    • Al-Sabi, M.N.S. et.al. (2022) [31270]
  • Sphyrna mokarran (Rüppell, 1837) (Carcharhiniformes: Sphyrnidae)
    • Al-Sabi, M.N.S. et.al. (2022) [31270]
Original description species: Al-Sabi, Ibrahim,Al-Hizab, Al-Jabr, Al-Shubaythi & Huffman, 2022
Genus: Huffmanela Moravec, 1987
Family: Trichosomoididae Yorke & Maplestone, 1926
Huffmanela sp. (2)

  • Carcharhinus dussumieri (Valenciennes in Müller & Henle, 1839) (Carcharhiniformes: Carcharhinidae)
    • Attia, M.M. et.al. (2021) [30038]
  • Carcharhinus plumbeus (Nardo, 1827) (Carcharhiniformes: Carcharhinidae)
    • Maclean, R.A. et.al. (2006) [9087]
Genus: Huffmanela Moravec, 1987
Family: Trichosomoididae Yorke & Maplestone, 1926
Hypanocotyle bullardi (4)
Chero, Cruces, Sáez, Camargo, Santos & Luque, 2018
  • Bathytoshia brevicaudata (Hutton, 1875) (Myliobatiformes: Dasyatidae)
    • Vaughan, D.B. et.al. (2023) [31765]
  • Hypanus dipterurus (Jordan & Gilbert, 1880) (Myliobatiformes: Dasyatidae)
    • Chero, J.D. et.al. (2018) [26182]
    • Chero De La Cruz, J.D. (2022) [32535]
    • Santillán, L.A. et.al. (2024) [33202]
Original description species: Chero, Cruces, Sáez, Camargo, Santos & Luque, 2018
Genus: Hypanocotyle Chero, Cruces, Sáez, Camargo, Santos & Luque, 2018
Family: Hexabothriidae Price, 1942
Order: Diclybothriidea
Subclass: Polyopisthocotylea
Hyperandrotrema cetorhini (4)
Maillard & Ktari, 1978
  • Cetorhinus maximus (Gunnerus, 1765) (Lamniformes: Cetorhinidae)
    • Maillard, C. et.al. (1978) [7849]
    • Bullard, S.A. et.al. (2006) [7400]
    • Orélis-Ribeiro, R. et.al. (2013) [19641]
    • Warren, M.B. et.al. (2019) [28146]
Original description species: Maillard & Ktari, 1978
Genus: Hyperandrotrema Maillard & Ktari, 1978
Family: Aporocotylidae Odhner, 1912
Superfamily: Schistosomatoidea
Suborder: Diplostomata
Order: Diplostomida
Subclass: Digenea
Hyperandrotrema walterboegeri (2)
Orélis-Ribeiro & Bullard, 2013
  • Isurus oxyrinchus Rafinesque, 1810 (Lamniformes: Lamnidae)
    • Orélis-Ribeiro, R. et.al. (2013) [19641]
    • Warren, M.B. et.al. (2019) [28146]
Original description species: Orélis-Ribeiro & Bullard, 2013
Genus: Hyperandrotrema Maillard & Ktari, 1978
Family: Aporocotylidae Odhner, 1912
Superfamily: Schistosomatoidea
Suborder: Diplostomata
Order: Diplostomida
Subclass: Digenea
Hysterothylacium aduncum (9)
(Rudolphi, 1802)
  • Bathyraja smirnovi (Soldatov & Pavlenko, 1915) (Rajiformes: Arhynchobatidae)
    • Peng, W.-F. et.al. (2011) [21235]
  • Cephaloscyllium isabellum (Bonnaterre, 1788) (Carcharhiniformes: Scyliorhinidae)
    • Hewitt, G.C. et.al. (1972) [16602] ( used parasite: Contracaecum (Thynnascaris) aduncum)
  • Dasyatis pastinaca (Linnaeus, 1758) (Myliobatiformes: Dasyatidae)
  • Etmopterus spinax (Linnaeus, 1758) (Squaliformes: Etmopteridae)
    • Klimpel, S. et.al. (2003) [5150]
  • Galeorhinus galeus (Linnaeus, 1758) (Carcharhiniformes: Triakidae)
    • Henderson, A.C. et.al. (2003) [9358]
  • Galeus melastomus Rafinesque, 1810 (Carcharhiniformes: Pentanchidae)
    • Dallares, S. et.al. (2017) [25922]
    • Dallares, S. et.al. (2017) [25639]
  • Raja clavata Linnaeus, 1758 (Rajiformes: Rajidae)
  • Rhinobatos rhinobatos (Linnaeus, 1758) (Rhinopristiformes: Rhinobatidae)
  • Scyliorhinus canicula (Linnaeus, 1758) (Carcharhiniformes: Scyliorhinidae)
    • Dallares, S. et.al. (2017) [25639]
  • Squalus acanthias Linnaeus, 1758 (Squaliformes: Squalidae)
    • Henderson, A.C. et.al. (2002) [9355]
    • Gaevskaya, A.V. (2012) [22626]
Synonyms: Ascaris adunca, Contracaecum (Thynnascaris) aduncum, Thynnascaris aduncum
Original description species: (Rudolphi, 1802)
Genus: Hysterothylacium Ward & Magath, 1917
Family: Anisakidae Railliet & Henry, 1912
Superfamily: Ascaridoidea
Order: Ascaridida
Hysterothylacium ganeshi (1)
Rajya Lakshmi & Sreeramulu, 2007
  • Eusphyra blochii (Cuvier, 1816) (Carcharhiniformes: Sphyrnidae)
    • Rajya Lakshmi, I. et.al. (2007) [21444] (used host: Sphyrna blochii )
Original description species: Rajya Lakshmi & Sreeramulu, 2007
Genus: Hysterothylacium Ward & Magath, 1917
Family: Anisakidae Railliet & Henry, 1912
Superfamily: Ascaridoidea
Order: Ascaridida
Hysterothylacium hospitum (1)
Solovjeva & Pozdnjakov, 1984
  • Prionace glauca (Linnaeus, 1758) (Carcharhiniformes: Carcharhinidae)
    • Bruce, N.L. et.al. (1994) [17029]
Original description species: Solovjeva & Pozdnjakov, 1984
Genus: Hysterothylacium Ward & Magath, 1917
Family: Anisakidae Railliet & Henry, 1912
Superfamily: Ascaridoidea
Order: Ascaridida
Hysterothylacium poecilurai (1)
Rajyalakshmi & Sreeramulu, 2005
  • Gymnura poecilura (Shaw, 1804) (Myliobatiformes: Gymnuridae)
    • Rajyalakshmi, I. et.al. (2005) [21461]
Original description species: Rajyalakshmi & Sreeramulu, 2005
Genus: Hysterothylacium Ward & Magath, 1917
Family: Anisakidae Railliet & Henry, 1912
Superfamily: Ascaridoidea
Order: Ascaridida
Hysterothylacium sp. (4)

  • Alopias vulpinus (Bonnaterre, 1788) (Lamniformes: Alopiidae)
  • Galeus melastomus Rafinesque, 1810 (Carcharhiniformes: Pentanchidae)
    • Dallares, S. et.al. (2017) [25922]
  • Pastinachus ater (Macleay, 1883) (Myliobatiformes: Dasyatidae)
    • Moravec, F. et.al. (2021) [29858]
  • Prionace glauca (Linnaeus, 1758) (Carcharhiniformes: Carcharhinidae)
  • Raja brachyura Lafont, 1873 (Rajiformes: Rajidae)
    • Alvarez, M.F. et.al. (2006) [9086]
  • Raja microocellata Montagu, 1818 (Rajiformes: Rajidae)
    • Alvarez, M.F. et.al. (2006) [9086]
Genus: Hysterothylacium Ward & Magath, 1917
Family: Anisakidae Railliet & Henry, 1912
Superfamily: Ascaridoidea
Order: Ascaridida